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Eldorado larkspur, Hansen's delphinium, Hansen's larkspur

dwarf, meadow, Nuttall's larkspur, or Sonne's larkspur, slim, thin-petal larkspur, two-lobe larkspur, upland larkspur

Stems

(25-)40-80(-180) cm;

base usually reddish, pubescent.

unbranched, 10-40(-70) cm;

base reddish, pubescence variable.

Leaves

blade pentagonal, 1.5-5 × 2.5-8 cm, long-pubescent, especially abaxially; ultimate lobes 0-18, width 4-20 mm (basal), 2-9 mm (cauline).

blade round, 1-6 × 2-12 cm, nearly glabrous; ultimate lobes 5-21, 5 or more extending more than 3/5 distance to petiole, width 1-7(-14) mm (basal), 0.5-6 mm (cauline), widest at middle or in proximal 1/2.

Inflorescences

(9-)15-40(-160)-flowered, dense to open;

pedicel 0.3-2.5(-6) cm, puberulent;

bracteoles 1-5(-8) mm from flowers, green, sometimes white-margined, linear-lanceolate, 2-6(-8) mm, puberulent.

4-18(-48)-flowered, at least 2 times as long as wide;

pedicel 0.8-6 cm, pubescence variable;

bracteoles 3-8(-18) mm from flowers, green to blue, linear, 3-7 mm, pubescence variable.

Flowers

sepals violet to white, ± puberulent, lateral sepals spreading to forward pointing, 7-10(-13) × 3-6(-8) mm, spurs gently upcurved, ascending 0-30° above horizontal, (6-)9-13(-16) mm;

lower petal blades elevated, ± exposing stamens, 3-7 mm, cleft 1-2(-4) mm;

hairs centered, densest on inner lobes near base of cleft, white.

sepals usually bluish purple, rarely white to pink, puberulent, lateral sepals reflexed or spreading, 8-21 × 3-10 mm, spurs decurved to straight, ascending 20-60° above horizontal, 8-23 mm;

lower petal blades elevated, exposing stamens, blue to purple, except sometimes in white-flowered plants, 4-11 mm, clefts 2-5 mm;

hairs mostly on inner lobes below junction of blade and claw, white, rarely yellow.

Fruits

8-20 mm, 2.2-4 times longer than wide, glabrous.

7-22 mm, 3.5-5 times longer than wide, glabrous to puberulent.

Seeds

echinate, appearing fuzzy to naked eye;

seed coat cells with margins straight, surfaces sparsely pustulate.

winged or not;

seed coat cell surfaces smooth or roughened, blunt hairs absent.

2n

= 16.

Delphinium hansenii

Delphinium nuttallianum

Phenology Flowering spring (-early summer).
Habitat Open coniferous woods, grassy sage scrub, meadow edges and well drained streamsides (generally not in very wet sites)
Elevation 300-3500 m (1000-11500 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Although Delphinium hansenii has often been confused with D. hesperium, seeds will instantly allow identification. Seeds of Delphinium hansenii are, as far as known, unique, bearing numerous, elongate, prismlike raised structures (extensions of single cells or small groups of cells) over the entire seed coat. If seeds are absent, larger flowers, more open inflorescences (except in D. hesperium subsp. cuyamacae), and general absence of pubescence of long hairs in D. hesperium are apparent upon comparison of the two species. Separating D. hansenii from D. variegatum may also be difficult. Again, seeds leave no doubt. In addition, smaller flowers and greater number of flowers per plant of D. hansenii should serve to distinguish D. hansenii from D. variegatum. White-flowered D. hansenii has been confused with D. gypsophilum and with D. hesperium subsp. pallescens. Other than seeds, pubescence of long hairs and smaller flowers present in D. hansenii and absent in the others will distinguish them.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium nuttallianum represents an extremely difficult complex, with many variations in a number of morphologic traits. The complex has been and continues to be a major source of confusion for identification of Delphinium in North America. Type specimens of D. nuttallianum represent plants growing under dry conditions in open areas. These are typically found at 1200-2000 m in sage scrub or lower montane forest. Delphinium nuttallianum may be confused with D. andersonii, D. antoninum, D. depauperatum, D. gracilentum, and two subspecies of D. patens (subsp. patens and subsp. montanum). Features that may be used to separate D. nuttallianum from the first four, are enumerated under the respective species discussions. From D. patens subsp. patens, D. nuttallianum may be distinguished by its narrower leaf lobes, larger fruits, and more compact inflorescence. The frequent presence of glandular hairs in the inflorescence of D. patens subsp. montanum, contrasted with their absence in D. nuttallianum, will separate these taxa. Dwarfed plants of D. polycladon may be confused with D. nuttallianum. The latter, however may be distinguished by its ringed seeds, and it does not have prominent buds or sigmoid pedicel.

Hybrids have been seen between Delphinium nuttallianum and D. andersonii, D. depauperatum (D. ×burkei Greene), D. distichum (D. ×diversicolor Rydberg), D. nudicaule, and D. polycladon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Sepals reddish purple to dark maroon.
subsp. ewanianum
1. Sepals dark blue-purple to white.
→ 2
2. Leaves primarily basal (although leaves may be dry at anthesis and thus lost in herbarium specimens, petiole base will be present); cauline leaves usually fewer than 3; sepals bright blue to white.
subsp. kernense
2. Leaves primarily cauline, basal leaves usually absent at anthesis; cauline leaves 3 or more; sepals dark blue-purple to white or pink.
subsp. hansenii
Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Echinata Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Subordinate taxa
D. hansenii subsp. ewanianum, D. hansenii subsp. hansenii, D. hansenii subsp. kernense
Synonyms D. hesperium var. hansenii D. pauciflorum, D. nuttallianum var. fulvum, D. nuttallianum var. levicaule, D. sonnei
Name authority (Greene) Greene: Pittonia 3: 94. (1896) Pritzel: in Walpers, Repert. Bot. Syst. 1: 744. (1842)
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