Delphinium hansenii |
Delphinium glaucum |
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Eldorado larkspur, Hansen's delphinium, Hansen's larkspur |
Brown's larkspur, duncecap larkspur, giant, giant larkspur, glaucous larkspur, Hooker's larkspur, mountain, mountain larkspur, or tall larkspur, pale larkspur, pale-flower Brown's larkspur, Sierra larkspur, tall larkspur, western larkspur |
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Stems | (25-)40-80(-180) cm; base usually reddish, pubescent. |
(60-)100-200(-300) cm; base usually green, glabrous, glaucous. |
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Leaves | blade pentagonal, 1.5-5 × 2.5-8 cm, long-pubescent, especially abaxially; ultimate lobes 0-18, width 4-20 mm (basal), 2-9 mm (cauline). |
blade round to pentagonal, 2-11 × 3-18 cm, margins seldom laciniate, glabrous; ultimate lobes 5-9(-15), width 5-24(-35) mm, tips abruptly tapered to mucronate apex; midcauline leaf lobes more than 3 times longer than wide. |
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Inflorescences | (9-)15-40(-160)-flowered, dense to open; pedicel 0.3-2.5(-6) cm, puberulent; bracteoles 1-5(-8) mm from flowers, green, sometimes white-margined, linear-lanceolate, 2-6(-8) mm, puberulent. |
(13-)40-90(-140)-flowered; pedicel 1-3(-5) cm, puberulent or glabrous; bracteoles 2-6(-10) mm from flowers, green to blue, linear, 2-7 mm, puberulent or glabrous. |
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Flowers | sepals violet to white, ± puberulent, lateral sepals spreading to forward pointing, 7-10(-13) × 3-6(-8) mm, spurs gently upcurved, ascending 0-30° above horizontal, (6-)9-13(-16) mm; lower petal blades elevated, ± exposing stamens, 3-7 mm, cleft 1-2(-4) mm; hairs centered, densest on inner lobes near base of cleft, white. |
sepals bluish purple to lavender, puberulent, lateral sepals forward pointing to spreading, 8-14(-21) × 3-6 mm, spurs straight, ascending to ca. 45°, 10-15(-19) mm; lower petal blades ± covering stamens, 4-6 mm, clefts 1-3 mm; hairs centered, mostly near base of cleft, white. |
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Fruits | 8-20 mm, 2.2-4 times longer than wide, glabrous. |
9-20 mm, 3.5-4.5 times longer than wide, glabrous to puberulent. |
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Seeds | echinate, appearing fuzzy to naked eye; seed coat cells with margins straight, surfaces sparsely pustulate. |
wing-margined; seed coat cells elongate but short, surfaces smooth or roughened. |
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2n | = 16. |
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Delphinium hansenii |
Delphinium glaucum |
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Phenology | Flowering summer. | |||||||||
Habitat | Meadows, wet thickets, bogs, streamsides, open coniferous woods | |||||||||
Elevation | 0-3200 m (0-10500 ft) | |||||||||
Distribution |
CA
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AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; MB; NT; SK; YT
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Discussion | Subspecies 3 (3 in the flora). Although Delphinium hansenii has often been confused with D. hesperium, seeds will instantly allow identification. Seeds of Delphinium hansenii are, as far as known, unique, bearing numerous, elongate, prismlike raised structures (extensions of single cells or small groups of cells) over the entire seed coat. If seeds are absent, larger flowers, more open inflorescences (except in D. hesperium subsp. cuyamacae), and general absence of pubescence of long hairs in D. hesperium are apparent upon comparison of the two species. Separating D. hansenii from D. variegatum may also be difficult. Again, seeds leave no doubt. In addition, smaller flowers and greater number of flowers per plant of D. hansenii should serve to distinguish D. hansenii from D. variegatum. White-flowered D. hansenii has been confused with D. gypsophilum and with D. hesperium subsp. pallescens. Other than seeds, pubescence of long hairs and smaller flowers present in D. hansenii and absent in the others will distinguish them. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
At the sites in Manitoba and Saskatchewan, Delphinium glaucum is naturalized, not native. Delphinium glaucum hybridizes extensively with D. barbeyi in Utah and Colorado to the extent that hybrids [D. ×occidentale (S. Watson) S. Watson] are more common in many areas than individuals of either parental stock. It occasionally hybridizes with D. distichum, D. polycladon, D. ramosum, and D. stachydeum. Hybrids with D. brachycentrum are called D. ×nutans A. Nelson. Tremendous variation is apparent in what is here recognized as Delphinium glaucum. This is the northern expression of the complex described in the discussion under Delphinium subsect. Exaltata. Although some geographic patterns are apparent in the variation within D. glaucum, infraspecific entities are not here recognized. Apparently because of rather recent and/or incomplete genetic isolation, the degree of differentiation between these units is not such that they can be consistently recognized. Specimens named Delphinium splendens represent plants grown in high-moisture, low-light conditions and may occur as sporadic individuals anywhere from California to Alaska. Type specimens of D. brownii Rydberg, D. canmorense Rydberg, and D. hookeri A. Nelson represent plants grown on relatively dry sites at high latitudes. Plants from dry sites at low latitudes are represented by D. bakerianum Bornmüller and D. occidentale var. reticulatum A. Nelson. Plants with lavender to white flowers are represented by type specimens of D. brownii forma pallidiflorum B. Boivin and D. cucullatum A. Nelson. Type specimens of D. alatum A. Nelson and D. glaucum var. alpinum F. L. Wynd (an invalid name) represent plants growing above or near treeline. Delphinium glaucum may be confused with D. californicum, D. exaltatum, D. polycladon, or D. stachydeum. For distinctions from D. californicum, see discussion under that species. Absence of basal or proximal cauline leaves, generally much larger plants (greater than 1.5 m), more flowers in the inflorescence, and shorter petioles on the leaves of D. glaucum are features that serve to distinguish this species from D. polycladon. In the latter, the leaves are primarily on the proximal stem, plants often less than 1.5 m, flowers more scattered, and petioles more than twice the length of leaf blades. Features of the sepals may be used to distinguish D. glaucum (dark lavender to blue purple, usually only minutely puberulent) from D. stachydeum (bright blue, densely puberulent). Vegetative parts of D. stachydeum are also densely puberulent, while those of D. glaucum typically are glabrous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3. | FNA vol. 3. | ||||||||
Parent taxa | Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Echinata | Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Exaltata | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | D. hesperium var. hansenii | D. scopulorum var. glaucum, D. splendens | ||||||||
Name authority | (Greene) Greene: Pittonia 3: 94. (1896) | S. Watson: Bot. California 2: 427. (1880) | ||||||||
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