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Eldorado larkspur, Hansen's delphinium, Hansen's larkspur

delphinium, larkspur

Habit Herbs, perennial, from fasciculate roots or rhizomes.
Stems

(25-)40-80(-180) cm;

base usually reddish, pubescent.

Leaves

blade pentagonal, 1.5-5 × 2.5-8 cm, long-pubescent, especially abaxially; ultimate lobes 0-18, width 4-20 mm (basal), 2-9 mm (cauline).

blade deeply palmately divided, round to pentagonal or reniform, margins entire or lobes apically crenate or lacerate, lobes of basal blades wider and fewer than those of cauline blades.

Inflorescences

(9-)15-40(-160)-flowered, dense to open;

pedicel 0.3-2.5(-6) cm, puberulent;

bracteoles 1-5(-8) mm from flowers, green, sometimes white-margined, linear-lanceolate, 2-6(-8) mm, puberulent.

terminal, 2-100(-more)-flowered racemes (occasionally branched, thus technically panicles), 5-40 cm or more;

bracts subtending inflorescence branches;

pedicels present or absent;

bracteoles (on pedicels) subopposite-subalternate, not forming involucre.

Flowers

sepals violet to white, ± puberulent, lateral sepals spreading to forward pointing, 7-10(-13) × 3-6(-8) mm, spurs gently upcurved, ascending 0-30° above horizontal, (6-)9-13(-16) mm;

lower petal blades elevated, ± exposing stamens, 3-7 mm, cleft 1-2(-4) mm;

hairs centered, densest on inner lobes near base of cleft, white.

bisexual, bilaterally symmetric;

sepals not persistent in fruit, 5;

upper sepal 1, spurred, 8-24 mm;

lateral sepals 2, ± ovate to elliptic, 8-18 mm;

lower sepals 2, similar to lateral sepals;

upper petals 2, spurred, enclosed in upper sepal, nectary inside tip of spur;

lower petals 2, plane, ± ovate, ± 2-lobed, clawed, 2-12 mm, nectary absent;

stamens 25-40;

filaments with base expanded;

staminodes absent between stamens and pistils;

pistils 3(-5), simple;

ovules 8-20 per pistil;

style present.

Fruits

8-20 mm, 2.2-4 times longer than wide, glabrous.

follicles, aggregate, sessile, ± curved-cylindric, sides prominently veined or not;

beak terminal, straight, 2-4 mm.

Seeds

echinate, appearing fuzzy to naked eye;

seed coat cells with margins straight, surfaces sparsely pustulate.

dark brown to black (often appearing white because of air in seed coat cells), rectangular to pyramidal, often ± rough surfaced.

x

= 8.

Delphinium hansenii

Delphinium

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from USDA
n temperate and arctic subtropical and; in Eastern Hemisphere; tropical mountains (s of equator in Africa)
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Although Delphinium hansenii has often been confused with D. hesperium, seeds will instantly allow identification. Seeds of Delphinium hansenii are, as far as known, unique, bearing numerous, elongate, prismlike raised structures (extensions of single cells or small groups of cells) over the entire seed coat. If seeds are absent, larger flowers, more open inflorescences (except in D. hesperium subsp. cuyamacae), and general absence of pubescence of long hairs in D. hesperium are apparent upon comparison of the two species. Separating D. hansenii from D. variegatum may also be difficult. Again, seeds leave no doubt. In addition, smaller flowers and greater number of flowers per plant of D. hansenii should serve to distinguish D. hansenii from D. variegatum. White-flowered D. hansenii has been confused with D. gypsophilum and with D. hesperium subsp. pallescens. Other than seeds, pubescence of long hairs and smaller flowers present in D. hansenii and absent in the others will distinguish them.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 300 (61 in the flora).

Three Eurasian species of Delphinium–D. elatum Linnaeus, D. grandiflorum Linnaeus, and D. tatsienense Franchet–have been commonly cultivated in North America. Of the nonnative taxa, only D. elatum is sporadically naturalized, as far as is known. Isolating mechanisms in Delphinium appear to be primarily ecological, geographic, and/or temporal. Where these distinctions are disrupted, introgression often exists. Hybridization occurs regularly between certain taxa, particularly in areas of disturbance (e.g., roadcuts, drainage ditches, clearcuts). The more common and easily recognized hybrids are included in the key.

Many names have been misapplied in Delphinium. The few misapplied names mentioned in discussions below refer to relatively widespread problems.

Unless otherwise noted, the key and descriptions refer to fresh material. Some features may be significantly altered by pressing; they can, however, usually be determined with a certain amount of effort and experience.

In the descriptions, "base of cleft" refers to the point where the cleft or sinus reaches most deeply into the petal blade.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Sepals reddish purple to dark maroon.
subsp. ewanianum
1. Sepals dark blue-purple to white.
→ 2
2. Leaves primarily basal (although leaves may be dry at anthesis and thus lost in herbarium specimens, petiole base will be present); cauline leaves usually fewer than 3; sepals bright blue to white.
subsp. kernense
2. Leaves primarily cauline, basal leaves usually absent at anthesis; cauline leaves 3 or more; sepals dark blue-purple to white or pink.
subsp. hansenii
1. Lower petal blades less than 1/5 length of lateral sepals; sepals never red or yellow.
Sect. Elatopsis
1. Lower petal blades more than 1/5 length of lateral sepals; sepals blue, purple, white, red, or yellow.
Sect. Diedropetala
Source FNA vol. 3. FNA vol. 3. Author: Michael J. Warnock.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Echinata Ranunculaceae
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Subordinate taxa
D. hansenii subsp. ewanianum, D. hansenii subsp. hansenii, D. hansenii subsp. kernense
Sect. Diedropetala, Sect. Elatopsis
Synonyms D. hesperium var. hansenii
Name authority (Greene) Greene: Pittonia 3: 94. (1896) Linnaeus: Sp. Pl. 1: 530. 175: Gen. Pl. ed 5, 236. (1754)
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