Delphinium gracilentum |
Ranunculaceae |
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Greene's larkspur, meadow larkspur, pine forest larkspur, slender or Greene's larkspur |
buttercup family, crowfoot family |
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Habit | Herbs, sometimes woody or herbaceous climbers or low shrubs, perennial or annual, often rhizomatous. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | (15-)30-50(-80) cm; base reddish, nearly glabrous. |
unarmed. |
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Leaves | blade round to pentagonal, 1.5-4 × 3-7 cm, nearly glabrous; ultimate lobes 3-7, distinctly wedge-shaped, usually 5 or fewer extending 3/5 distance to petiole, width 5-20 mm (basal), 1-15 mm (cauline), widest in distal 1/2. |
blade undivided or more commonly divided or compound, base cordate, sometimes truncate or cuneate, margins entire, toothed, or incised; venation pinnate or palmate. |
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Inflorescences | 5-20(-38)-flowered; pedicel spreading from rachis at nearly 90°, 1-3(-4) cm, glabrous or glandular-pubescent; bracteoles (7-)11-19 mm from flowers, blue or green, linear, 2-5 mm, puberulent to glabrous. |
terminal or axillary, racemes, cymes, umbels, panicles, or spikes, or flowers solitary, flowers pedicellate or sessile. |
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Flowers | sepals dark bluish purple to pink or white, usually retaining color upon drying, glabrous, lateral sepals reflexed, 6-10(-13) × 3-6 mm, spurs often curved upward, within 30° above or below horizontal, 8-12(-14) mm; lower petal blades elevated, exposing stamens, 3-5 mm, clefts 1-3 mm; hairs almost exclusively near base of cleft, centered or mostly on inner lobes, usually yellow. |
bisexual, sometimes unisexual, inconspicuous or showy, radially or bilaterally symmetric; sepaloid bracteoles absent; perianth hypogynous; sepals usually imbricate, 3-6(-20), distinct, often petaloid and colored, occasionally spurred; petals 0-26, distinct (connate in Consolida), plane, cup-shaped, funnel-shaped, or spurred, conspicuous or greatly reduced; nectary usually present, rarely absent; stamens 5-many, distinct; anthers dehiscing longitudinally; staminodes absent (except in Aquilegia and Clematis); pistils 1-many; styles present or absent, often persistent in fruit as beak. |
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Fruits | 8-16 mm, 3-3.5 times longer than wide, glabrous to glandular-puberulent. |
achenes, follicles, or rarely utricles, capsules, or berries, often aggregated into globose to cylindric heads. |
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Seeds | unwinged; seed coats ± pitted, cell surfaces roughened. |
1-many per ovary, never stalked, not arillate; endosperm abundant; embryo usually small. |
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Delphinium gracilentum |
Ranunculaceae |
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Phenology | Flowering spring–early summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Open coniferous forest | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 150-2700 m (500-8900 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA
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Worldwide |
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Discussion | Delphinium gracilentum hybridizes with D. patens subsp. patens in the northern Sierra Nevada foothills and is very similar to that species, making hybrids difficult to discern. While D. gracilentum and D. patens are easily distinguished in most of their ranges, morphologic distinctions between the two taxa are blurred in the northern Sierra Nevada foothills region, particularly in Butte County, California. Coniferous woods are preferred by D. gracilentum; D. patens subspp. patens and hepaticoideum are more often found in broadleaf woods. The former species has more widely spreading pedicels than the latter, and D. gracilentum usually has wider leaf lobes than D. patens subsp. patens. In the southern Sierra Nevada, D. gracilentum may come in contact with D. patens subsp. montanum. Though hybrids are not common, some gene flow has apparently occurred. Sepal color phases are not stable and considerable variation occurs within populations. The type specimen of Delphinium gracilentum represents the northern, lower elevation, nonglandular, dark-flowered phase. The type specimen of D. gracilentum forma versicolor Ewan differs only by its pink or white flowers. A limited range of intermediate colors occurs, and populations may be made up of plants of a single color or several different colors. The type specimen of D. greenei Eastwood represents the southern, higher elevation, glandular (at least on pedicels) expression. The type specimen includes representatives of dark- and light-flowered individuals of this phase. The type specimen of D. gracilentum forma versicolor (not seen by the author) is the "albino" phase referred to by Greene in his description of D. gracilentum. Several of the paratypes cited by Ewan have been seen, as have a number of individuals in natural populations. Delphinium gracilentum has been confused with D. patens or D. nuttallianum. Delphinium gracilentum may be distinguished from D. nuttallianum by its wider leaf lobes, smaller fruits, and more elongate inflorescences, and from D. patens by its wider leaf lobes, more open inflorescences, and usually shorter fruits. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera ca. 60, species 1700 (22 genera, 284 species in the flora). The flowers of many species of Ranunculaceae begin to open long before anthesis, while the floral organs are just partly expanded. Only mature flowers with open anthers should be used for determination of diagnostic characteristics (especially measurements). The literature is inconsistent about the term for the whorl of organs between sepals and stamens; these may be conspicuous and petaloid, or reduced to stalked nectaries, or intermediate between the two states. They have been called petals, honey-leaves, or (when they are inconspicuous) staminodes or nectaries. We follow M. Tamura (1993) and treat as petals all organs between the sepals and stamens, except in Clematis and Aquilegia where they usually bear rudimentary anthers and clearly represent staminodes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3. | FNA vol. 3, p. 85. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | D. patens subsp. greenei, D. pratense | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Greene: Pittonia 3: 15. (1896) | Jussieu | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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