Flora of North America species comparison

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Delphinium gracilentum

Delphinium inopinum

Greene's larkspur, meadow larkspur, pine forest larkspur, slender or Greene's larkspur

unexpected larkspur
Stems

(15-)30-50(-80) cm;

base reddish, nearly glabrous.

70-110(-150) cm;

base reddish or not, glabrous, often glaucous.
Leaves

blade round to pentagonal, 1.5-4 × 3-7 cm, nearly glabrous; ultimate lobes 3-7, distinctly wedge-shaped, usually 5 or fewer extending 3/5 distance to petiole, width 5-20 mm (basal), 1-15 mm (cauline), widest in distal 1/2.

blade ± pentagonal, 1-5 × 1.5-7 cm, glabrous; ultimate lobes 3-9, width 5-28 mm (basal), 3-18 mm (cauline).
Inflorescences

5-20(-38)-flowered;

pedicel spreading from rachis at nearly 90°, 1-3(-4) cm, glabrous or glandular-pubescent;

bracteoles (7-)11-19 mm from flowers, blue or green, linear, 2-5 mm, puberulent to glabrous.

20-35(-51)-flowered, usually dense;

pedicel 0.3-1.5(-2.5) cm, glabrous;

bracteoles 2-4 mm from flowers, green, linear, 1-2(-4) mm, nearly glabrous.
Flowers

sepals dark bluish purple to pink or white, usually retaining color upon drying, glabrous, lateral sepals reflexed, 6-10(-13) × 3-6 mm, spurs often curved upward, within 30° above or below horizontal, 8-12(-14) mm;

lower petal blades elevated, exposing stamens, 3-5 mm, clefts 1-3 mm;

hairs almost exclusively near base of cleft, centered or mostly on inner lobes, usually yellow.

sepals white to light blue, glabrous, lateral sepals spreading to forward pointing, 8-12 × 3-5 mm, spurs straight to gently upcurved, ascending 30-60° above horizontal, 9-12 mm;

lower petal blades slightly elevated, ± exposing stamens, 3-5 mm, clefts 1-2 mm;

hairs centered, densest near base of cleft, white.
Fruits

8-16 mm, 3-3.5 times longer than wide, glabrous to glandular-puberulent.

12-20 mm, 2.6-4 times longer than wide, glabrous.
Seeds

unwinged;

seed coats ± pitted, cell surfaces roughened.

wing-margined;

seed coat cells surfaces smooth.

Delphinium gracilentum

Delphinium inopinum
Phenology Flowering spring–early summer. Flowering summer.
Habitat Open coniferous forest Rock outcrops in open coniferous woods
Elevation 150-2700 m (500-8900 ft) 2200-2800 m (7200-9200 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Delphinium gracilentum hybridizes with D. patens subsp. patens in the northern Sierra Nevada foothills and is very similar to that species, making hybrids difficult to discern. While D. gracilentum and D. patens are easily distinguished in most of their ranges, morphologic distinctions between the two taxa are blurred in the northern Sierra Nevada foothills region, particularly in Butte County, California. Coniferous woods are preferred by D. gracilentum; D. patens subspp. patens and hepaticoideum are more often found in broadleaf woods. The former species has more widely spreading pedicels than the latter, and D. gracilentum usually has wider leaf lobes than D. patens subsp. patens. In the southern Sierra Nevada, D. gracilentum may come in contact with D. patens subsp. montanum. Though hybrids are not common, some gene flow has apparently occurred.

Sepal color phases are not stable and considerable variation occurs within populations. The type specimen of Delphinium gracilentum represents the northern, lower elevation, nonglandular, dark-flowered phase. The type specimen of D. gracilentum forma versicolor Ewan differs only by its pink or white flowers. A limited range of intermediate colors occurs, and populations may be made up of plants of a single color or several different colors. The type specimen of D. greenei Eastwood represents the southern, higher elevation, glandular (at least on pedicels) expression. The type specimen includes representatives of dark- and light-flowered individuals of this phase. The type specimen of D. gracilentum forma versicolor (not seen by the author) is the "albino" phase referred to by Greene in his description of D. gracilentum. Several of the paratypes cited by Ewan have been seen, as have a number of individuals in natural populations.

Delphinium gracilentum has been confused with D. patens or D. nuttallianum. Delphinium gracilentum may be distinguished from D. nuttallianum by its wider leaf lobes, smaller fruits, and more elongate inflorescences, and from D. patens by its wider leaf lobes, more open inflorescences, and usually shorter fruits.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium inopinum is apparently endemic to a white metamorphic rock substrate in the Piute Mountains and southern Sierra Nevada. It is not known to hybridize with any other species, although D. patens subsp. montanum has been collected (when both were flowering) within 1 km of D. inopinum and probably occurs much closer. Delphinium inopinum is often confused with D. parishii subsp. pallidum and superficially resembles some white-flowered individuals of D. hansenii, as well as D. gypsophilum and D. hesperium subsp. pallescens. The massive roots with prominent buds readily distinguish D. inopinum from all of these. In addition, the pubescence found on D. hansenii will separate it from the glabrous D. inopinum. Leaves are rarely seen at anthesis near the base of the stem in D. hesperium subsp. pallescens; they are present in D. inopinum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms D. patens subsp. greenei, D. pratense D. parishii var. inopinum
Name authority Greene: Pittonia 3: 15. (1896) (Jepson) H. F. Lewis & Epling: Brittonia 8: 11. (1954)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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