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Carolina larkspur

dwarf, meadow, Nuttall's larkspur, or Sonne's larkspur, slim, thin-petal larkspur, two-lobe larkspur, upland larkspur

Stems

(20-)40-90(-150) cm;

base reddish or not, ± pubescent.

unbranched, 10-40(-70) cm;

base reddish, pubescence variable.

Leaves

blade round to pentagonal, 1-8 × 2-12 cm, pubescence variable; ultimate lobes 3-29, width 2-10 mm (basal), 1-7 mm (cauline).

blade round, 1-6 × 2-12 cm, nearly glabrous; ultimate lobes 5-21, 5 or more extending more than 3/5 distance to petiole, width 1-7(-14) mm (basal), 0.5-6 mm (cauline), widest at middle or in proximal 1/2.

Inflorescences

(3-)8-27(-94)-flowered;

pedicel (0.4-)0.7-1.8(-5.7) cm, nearly glabrous to glandular;

bracteoles 1-3.5(-6) mm from flowers, green or blue, linear, 2-7 mm, pubescence nearly glabrous to glandular.

4-18(-48)-flowered, at least 2 times as long as wide;

pedicel 0.8-6 cm, pubescence variable;

bracteoles 3-8(-18) mm from flowers, green to blue, linear, 3-7 mm, pubescence variable.

Flowers

sepals purple to blue to white, nearly glabrous, lateral sepals spreading, (7-)9-14(-17) × (3-)3.5-6(-8) mm, spurs ± upcurved, ascending 20-90° above vertical, (9-)11-17(-19) mm;

lower petal blades ± covering stamens, 5-7 mm, cleft 2-4 mm;

hairs centered, densest near base of cleft, white, sometimes blue or yellow.

sepals usually bluish purple, rarely white to pink, puberulent, lateral sepals reflexed or spreading, 8-21 × 3-10 mm, spurs decurved to straight, ascending 20-60° above horizontal, 8-23 mm;

lower petal blades elevated, exposing stamens, blue to purple, except sometimes in white-flowered plants, 4-11 mm, clefts 2-5 mm;

hairs mostly on inner lobes below junction of blade and claw, white, rarely yellow.

Fruits

(10-)12.5-18.5(-27) mm, 4-4.5 times longer than wide, glabrous to puberulent.

7-22 mm, 3.5-5 times longer than wide, glabrous to puberulent.

Seeds

seed coat cells with surfaces pustulate or smooth.

winged or not;

seed coat cell surfaces smooth or roughened, blunt hairs absent.

2n

= 16.

Delphinium carolinianum

Delphinium nuttallianum

Phenology Flowering spring (-early summer).
Habitat Open coniferous woods, grassy sage scrub, meadow edges and well drained streamsides (generally not in very wet sites)
Elevation 300-3500 m (1000-11500 ft)
Distribution
from FNA
AL; AR; CO; FL; GA; IA; IL; KS; KY; LA; MN; MO; MS; NC; ND; NE; OK; SC; SD; TN; TX; WI; MB; Mexico
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from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
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Discussion

Subspecies 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium nuttallianum represents an extremely difficult complex, with many variations in a number of morphologic traits. The complex has been and continues to be a major source of confusion for identification of Delphinium in North America. Type specimens of D. nuttallianum represent plants growing under dry conditions in open areas. These are typically found at 1200-2000 m in sage scrub or lower montane forest. Delphinium nuttallianum may be confused with D. andersonii, D. antoninum, D. depauperatum, D. gracilentum, and two subspecies of D. patens (subsp. patens and subsp. montanum). Features that may be used to separate D. nuttallianum from the first four, are enumerated under the respective species discussions. From D. patens subsp. patens, D. nuttallianum may be distinguished by its narrower leaf lobes, larger fruits, and more compact inflorescence. The frequent presence of glandular hairs in the inflorescence of D. patens subsp. montanum, contrasted with their absence in D. nuttallianum, will separate these taxa. Dwarfed plants of D. polycladon may be confused with D. nuttallianum. The latter, however may be distinguished by its ringed seeds, and it does not have prominent buds or sigmoid pedicel.

Hybrids have been seen between Delphinium nuttallianum and D. andersonii, D. depauperatum (D. ×burkei Greene), D. distichum (D. ×diversicolor Rydberg), D. nudicaule, and D. polycladon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves absent at anthesis, cauline leaves divided into many narrow (less than 2 mm wide) segments, blade not distinctly 3-parted; distalmost petiole less than 5 mm; sepals usually blue or purple (rarely white).
subsp. carolinianum
1. Basal leaves usually present at anthesis, and/or cauline leaf lobes usually wider than 2 mm, blade often distinctly 3-parted or more; distalmost petiole more than 5 mm; sepals blue or white.
→ 2
2. Leaf blade distinctly 3-parted with few additional divisions; sepals blue to white; roots usually ± vertical, often without major branches.
subsp. vimineum
2. Leaf blade with 3–5 or more major divisions, each further divided into segments; sepals white to very pale blue; roots ±horizontal with several major branches.
→ 3
3. Stems usually less than 45 cm; in thin soils over limestone in clearings of deciduous woods; leaf blade with 3 major divisions; e of Mississippi River.
subsp. calciphilum
3. Stems usually more than 45 cm; in deeper soils in grasslands; leaf blade with 5 or more major divisions; w of Mississippi River.
subsp. virescens
Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Virescens Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Subordinate taxa
D. carolinianum subsp. calciphilum, D. carolinianum subsp. carolinianum, D. carolinianum subsp. vimineum, D. carolinianum subsp. virescens
Synonyms D. pauciflorum, D. nuttallianum var. fulvum, D. nuttallianum var. levicaule, D. sonnei
Name authority Walter: Fl. Carol., 155. (1788) Pritzel: in Walpers, Repert. Bot. Syst. 1: 744. (1842)
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