FNA: Delphinium cardinale vs. Delphinium newtonianum

	Delphinium cardinale	Delphinium newtonianum
	cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur	newton's larkspur, Ozark larkspur
Stems	(33-)50-150(-280) cm; base reddish, ± puberulent.	40-90 cm; base often reddish, puberulent.
Leaves	blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).	blade round to pentagonal, 4-7 × 5-15 cm, nearly glabrous; ultimate lobes 3-7, width 8-20 mm (basal), 5-15 mm (cauline), widest at middle or in proximal 1/2.
Inflorescences	10-40(-80)-flowered, open, narrowly pyramidal; pedicel spreading, (1-)2-5 cm, ± puberulent; bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.	8-20(-40)-flowered, as wide as long or nearly so; pedicel 1-4(-6) cm, pubescent; bracteoles 6-15 mm from flowers, green, linear, 1.5-5 mm, puberulent.
Flowers	sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm; lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm; hairs centered at base of cleft, short, sparse, yellow.	sepals dark to light blue, rarely white, glabrous, lateral sepals spreading, 12-14 × 6-7 mm, spurs straight to decurved, within 30° of horizontal, 10-15 mm; lower petal blades slightly elevated, ± exposing stamens, 4-5 mm, clefts 2-3 mm; hairs mostly centered near base of cleft, yellow.
Fruits	erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.	8-12 mm, 3-3.5 times longer than wide, nearly glabrous.
Seeds	unwinged; seed coat cells with margins undulate, surfaces roughened.	unwinged; surface of each seed coat cell with swollen, blunt, hairlike structures, barely visible at 20x, otherwise smooth.
2n	= 16.

	Delphinium cardinale	Delphinium newtonianum
Phenology	Flowering spring–early summer.	Flowering early summer.
Habitat	Slopes (often unstable) in chaparral	Slopes in deciduous forest
Elevation	50-1500 m (200-4900 ft)	500-700 m (1600-2300 ft)
Distribution	CA; Mexico (Baja California, Baja California Sur) [WildflowerSearch map] [BONAP county map]	AR [BONAP county map]
Discussion	Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild. Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study. The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.)	No cases of hybridization are known. Delphinium newtonianum often occurs in mixed populations with D. tricorne. It normally does not begin flowering until 4-6 weeks after D. tricorne has finished. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Name authority	Hooker: Bot. Mag., plate 4887. (1855)	D. M. Moore: Rhodora 41: 196. (1939)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium newtonianum

cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur

newton's larkspur, Ozark larkspur

Stems

(33-)50-150(-280) cm;

base reddish, ± puberulent.

40-90 cm;

base often reddish, puberulent.

Leaves

blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).

blade round to pentagonal, 4-7 × 5-15 cm, nearly glabrous; ultimate lobes 3-7, width 8-20 mm (basal), 5-15 mm (cauline), widest at middle or in proximal 1/2.

Inflorescences

10-40(-80)-flowered, open, narrowly pyramidal;

pedicel spreading, (1-)2-5 cm, ± puberulent;

bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.

8-20(-40)-flowered, as wide as long or nearly so;

pedicel 1-4(-6) cm, pubescent;

bracteoles 6-15 mm from flowers, green, linear, 1.5-5 mm, puberulent.

Flowers

sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm;

lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm;

hairs centered at base of cleft, short, sparse, yellow.

sepals dark to light blue, rarely white, glabrous, lateral sepals spreading, 12-14 × 6-7 mm, spurs straight to decurved, within 30° of horizontal, 10-15 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-5 mm, clefts 2-3 mm;

hairs mostly centered near base of cleft, yellow.

Fruits

erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.

8-12 mm, 3-3.5 times longer than wide, nearly glabrous.

Seeds

unwinged;

seed coat cells with margins undulate, surfaces roughened.

unwinged;

surface of each seed coat cell with swollen, blunt, hairlike structures, barely visible at 20x, otherwise smooth.

= 16.

Delphinium newtonianum

Phenology

Flowering spring–early summer.

Flowering early summer.

Habitat

Slopes (often unstable) in chaparral

Slopes in deciduous forest

Elevation

50-1500 m (200-4900 ft)

500-700 m (1600-2300 ft)

Distribution

CA; Mexico (Baja California, Baja California Sur)

[WildflowerSearch map]

[BONAP county map]

Discussion

Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild.

Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study.

The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

No cases of hybridization are known. Delphinium newtonianum often occurs in mixed populations with D. tricorne. It normally does not begin flowering until 4-6 weeks after D. tricorne has finished.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Name authority

Hooker: Bot. Mag., plate 4887. (1855)

D. M. Moore: Rhodora 41: 196. (1939)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Delphinium cardinale

Delphinium newtonianum

Delphinium cardinale

Delphinium newtonianum