FNA: Delphinium cardinale vs. Delphinium inopinum

	Delphinium cardinale	Delphinium inopinum
	cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur	unexpected larkspur
Stems	(33-)50-150(-280) cm; base reddish, ± puberulent.	70-110(-150) cm; base reddish or not, glabrous, often glaucous.
Leaves	blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).	blade ± pentagonal, 1-5 × 1.5-7 cm, glabrous; ultimate lobes 3-9, width 5-28 mm (basal), 3-18 mm (cauline).
Inflorescences	10-40(-80)-flowered, open, narrowly pyramidal; pedicel spreading, (1-)2-5 cm, ± puberulent; bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.	20-35(-51)-flowered, usually dense; pedicel 0.3-1.5(-2.5) cm, glabrous; bracteoles 2-4 mm from flowers, green, linear, 1-2(-4) mm, nearly glabrous.
Flowers	sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm; lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm; hairs centered at base of cleft, short, sparse, yellow.	sepals white to light blue, glabrous, lateral sepals spreading to forward pointing, 8-12 × 3-5 mm, spurs straight to gently upcurved, ascending 30-60° above horizontal, 9-12 mm; lower petal blades slightly elevated, ± exposing stamens, 3-5 mm, clefts 1-2 mm; hairs centered, densest near base of cleft, white.
Fruits	erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.	12-20 mm, 2.6-4 times longer than wide, glabrous.
Seeds	unwinged; seed coat cells with margins undulate, surfaces roughened.	wing-margined; seed coat cells surfaces smooth.
2n	= 16.

	Delphinium cardinale	Delphinium inopinum
Phenology	Flowering spring–early summer.	Flowering summer.
Habitat	Slopes (often unstable) in chaparral	Rock outcrops in open coniferous woods
Elevation	50-1500 m (200-4900 ft)	2200-2800 m (7200-9200 ft)
Distribution	CA; Mexico (Baja California, Baja California Sur) [WildflowerSearch map] [BONAP county map]	CA [BONAP county map]
Discussion	Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild. Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study. The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium inopinum is apparently endemic to a white metamorphic rock substrate in the Piute Mountains and southern Sierra Nevada. It is not known to hybridize with any other species, although D. patens subsp. montanum has been collected (when both were flowering) within 1 km of D. inopinum and probably occurs much closer. Delphinium inopinum is often confused with D. parishii subsp. pallidum and superficially resembles some white-flowered individuals of D. hansenii, as well as D. gypsophilum and D. hesperium subsp. pallescens. The massive roots with prominent buds readily distinguish D. inopinum from all of these. In addition, the pubescence found on D. hansenii will separate it from the glabrous D. inopinum. Leaves are rarely seen at anthesis near the base of the stem in D. hesperium subsp. pallescens; they are present in D. inopinum. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. parishii var. inopinum
Name authority	Hooker: Bot. Mag., plate 4887. (1855)	(Jepson) H. F. Lewis & Epling: Brittonia 8: 11. (1954)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium inopinum

cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur

unexpected larkspur

Stems

(33-)50-150(-280) cm;

base reddish, ± puberulent.

70-110(-150) cm;

base reddish or not, glabrous, often glaucous.

Leaves

blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).

blade ± pentagonal, 1-5 × 1.5-7 cm, glabrous; ultimate lobes 3-9, width 5-28 mm (basal), 3-18 mm (cauline).

Inflorescences

10-40(-80)-flowered, open, narrowly pyramidal;

pedicel spreading, (1-)2-5 cm, ± puberulent;

bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.

20-35(-51)-flowered, usually dense;

pedicel 0.3-1.5(-2.5) cm, glabrous;

bracteoles 2-4 mm from flowers, green, linear, 1-2(-4) mm, nearly glabrous.

Flowers

sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm;

lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm;

hairs centered at base of cleft, short, sparse, yellow.

sepals white to light blue, glabrous, lateral sepals spreading to forward pointing, 8-12 × 3-5 mm, spurs straight to gently upcurved, ascending 30-60° above horizontal, 9-12 mm;

lower petal blades slightly elevated, ± exposing stamens, 3-5 mm, clefts 1-2 mm;

hairs centered, densest near base of cleft, white.

Fruits

erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.

12-20 mm, 2.6-4 times longer than wide, glabrous.

Seeds

unwinged;

seed coat cells with margins undulate, surfaces roughened.

wing-margined;

seed coat cells surfaces smooth.

= 16.

Delphinium inopinum

Phenology

Flowering spring–early summer.

Flowering summer.

Habitat

Slopes (often unstable) in chaparral

Rock outcrops in open coniferous woods

Elevation

50-1500 m (200-4900 ft)

2200-2800 m (7200-9200 ft)

Distribution

CA; Mexico (Baja California, Baja California Sur)

[WildflowerSearch map]

[BONAP county map]

Discussion

Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild.

Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study.

The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium inopinum is apparently endemic to a white metamorphic rock substrate in the Piute Mountains and southern Sierra Nevada. It is not known to hybridize with any other species, although D. patens subsp. montanum has been collected (when both were flowering) within 1 km of D. inopinum and probably occurs much closer. Delphinium inopinum is often confused with D. parishii subsp. pallidum and superficially resembles some white-flowered individuals of D. hansenii, as well as D. gypsophilum and D. hesperium subsp. pallescens. The massive roots with prominent buds readily distinguish D. inopinum from all of these. In addition, the pubescence found on D. hansenii will separate it from the glabrous D. inopinum. Leaves are rarely seen at anthesis near the base of the stem in D. hesperium subsp. pallescens; they are present in D. inopinum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. parishii var. inopinum

Name authority

Hooker: Bot. Mag., plate 4887. (1855)

(Jepson) H. F. Lewis & Epling: Brittonia 8: 11. (1954)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Delphinium cardinale

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