FNA: Delphinium cardinale vs. Delphinium geraniifolium

	Delphinium cardinale	Delphinium geraniifolium
	cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur	Clark Valley larkspur, Mogollon larkspur
Stems	(33-)50-150(-280) cm; base reddish, ± puberulent.	60-100 cm; base reddish, puberulent, midstems pubescent.
Leaves	blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).	blade dark green, at least adaxially, fan-shaped to reniform, 2-5 × 3-7 cm, pubescent, especially abaxially; ultimate lobes 5-15, width 4-15 mm (basal lobes 5-15 mm), apex abruptly tapered, usually mucronate; veins prominent.
Inflorescences	10-40(-80)-flowered, open, narrowly pyramidal; pedicel spreading, (1-)2-5 cm, ± puberulent; bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.	20-90-flowered, dense, cylindric; pedicel ascending to spreading, 1-2 cm, puberulent; bracteoles 1-2 mm from flowers, green, linear, 4-6 mm, puberulent.
Flowers	sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm; lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm; hairs centered at base of cleft, short, sparse, yellow.	sepals dark blue to purple, puberulent, lateral sepals spreading to slightly forward pointing, 10-14 × 3-5 mm, spurs ascending 20-70°, truncate or downcurved apically, 12-15 mm; lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 0.5-2 mm; hairs sparse, local below junction of blade and claw, scattered on margins, white.
Fruits	erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.	13-18 mm, 3-3.5 times longer than wide, puberulent.
Seeds	unwinged; seed coat cells with margins undulate, surfaces roughened.	unwinged; seed coat cells with margins straight, surfaces ± roughened.
2n	= 16.	= 16.

	Delphinium cardinale	Delphinium geraniifolium
Phenology	Flowering spring–early summer.	Flowering summer.
Habitat	Slopes (often unstable) in chaparral	Heavy clay soil, dry meadows in coniferous woods
Elevation	50-1500 m (200-4900 ft)	1800-3400 m (5900-11200 ft)
Distribution	CA; Mexico (Baja California, Baja California Sur) [WildflowerSearch map] [BONAP county map]	AZ; NM [WildflowerSearch map] [BONAP county map]
Discussion	Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild. Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study. The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium geraniifolium is a more pubescent analog of the closely related D. scopulorum, the former occurring in heavier soils at higher elvation. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. tenuisectum subsp. amplibracteatum
Name authority	Hooker: Bot. Mag., plate 4887. (1855)	Rydberg: Bull. Torrey Bot. Club 26: 583. (1899)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium geraniifolium

cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur

Clark Valley larkspur, Mogollon larkspur

Stems

(33-)50-150(-280) cm;

base reddish, ± puberulent.

60-100 cm;

base reddish, puberulent, midstems pubescent.

Leaves

blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).

blade dark green, at least adaxially, fan-shaped to reniform, 2-5 × 3-7 cm, pubescent, especially abaxially; ultimate lobes 5-15, width 4-15 mm (basal lobes 5-15 mm), apex abruptly tapered, usually mucronate;

veins prominent.

Inflorescences

10-40(-80)-flowered, open, narrowly pyramidal;

pedicel spreading, (1-)2-5 cm, ± puberulent;

bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.

20-90-flowered, dense, cylindric;

pedicel ascending to spreading, 1-2 cm, puberulent;

bracteoles 1-2 mm from flowers, green, linear, 4-6 mm, puberulent.

Flowers

sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm;

lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm;

hairs centered at base of cleft, short, sparse, yellow.

sepals dark blue to purple, puberulent, lateral sepals spreading to slightly forward pointing, 10-14 × 3-5 mm, spurs ascending 20-70°, truncate or downcurved apically, 12-15 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 0.5-2 mm;

hairs sparse, local below junction of blade and claw, scattered on margins, white.

Fruits

erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.

13-18 mm, 3-3.5 times longer than wide, puberulent.

Seeds

unwinged;

seed coat cells with margins undulate, surfaces roughened.

unwinged;

seed coat cells with margins straight, surfaces ± roughened.

= 16.

Delphinium cardinale

Delphinium geraniifolium

Phenology

Flowering spring–early summer.

Flowering summer.

Habitat

Slopes (often unstable) in chaparral

Heavy clay soil, dry meadows in coniferous woods

Elevation

50-1500 m (200-4900 ft)

1800-3400 m (5900-11200 ft)

Distribution

CA; Mexico (Baja California, Baja California Sur)

[WildflowerSearch map]

[BONAP county map]

AZ; NM

[WildflowerSearch map]

[BONAP county map]

Discussion

Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild.

Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study.

The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium geraniifolium is a more pubescent analog of the closely related D. scopulorum, the former occurring in heavier soils at higher elvation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. tenuisectum subsp. amplibracteatum

Name authority

Hooker: Bot. Mag., plate 4887. (1855)

Rydberg: Bull. Torrey Bot. Club 26: 583. (1899)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Delphinium cardinale

Delphinium geraniifolium

Delphinium cardinale

Delphinium geraniifolium