FNA: Delphinium cardinale vs. Delphinium andesicola

	Delphinium cardinale	Delphinium andesicola
	cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur	Chiricahua Mountain larkspur
Stems	(33-)50-150(-280) cm; base reddish, ± puberulent.	60-200 +cm; base reddish or not, glabrous, glaucous.
Leaves	blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).	blade cordate to semicircular, 5-8 × 5-12 cm, nearly glabrous; ultimate lobes 5-16, width 3-20 mm, tips gradually tapered to mucronate apex; midcauline leaf lobes more than 3 times longer than wide.
Inflorescences	10-40(-80)-flowered, open, narrowly pyramidal; pedicel spreading, (1-)2-5 cm, ± puberulent; bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.	20-80-flowered; pedicel 1-2(-3) cm, puberulent; bracteoles 1-3 mm from flowers, green to brown, linear-lanceolate, 3-6 mm, puberulent.
Flowers	sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm; lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm; hairs centered at base of cleft, short, sparse, yellow.	sepals purple, puberulent, lateral sepals spreading, 9-12 × 5-7 mm, spurs ascending ca. 45°, curved downward apically, purple, 10-13 mm, blunt tipped; lower petal blades ± covering stamens, 4-6 mm, clefts 1.5-2.5 mm; hairs centered, densest on inner lobes near base of cleft, white.
Fruits	erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.	12-15 mm, 3.5-4 times longer than wide, sparsely puberulent.
Seeds	unwinged; seed coat cells with margins undulate, surfaces roughened.	unwinged; seed coat cells elongate, surfaces pustulate.
2n	= 16.	= 16.

	Delphinium cardinale	Delphinium andesicola
Phenology	Flowering spring–early summer.	Flowering summer–early fall.
Habitat	Slopes (often unstable) in chaparral	Meadows and coniferous woods
Elevation	50-1500 m (200-4900 ft)	2200-3200 m (7200-10500 ft)
Distribution	CA; Mexico (Baja California, Baja California Sur) [WildflowerSearch map] [BONAP county map]	AZ [BONAP county map]
Discussion	Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild. Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study. The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium andesicola, the westernmost representative of the southern Cordilleran complex, is found in the Chiricahua, Huachuca, Graham, and White mountains. Hybrids with Delphinium scopulorum are known. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Exaltata
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. andesicola subsp. amplum
Name authority	Hooker: Bot. Mag., plate 4887. (1855)	Ewan: J. Washington Acad. Sci. 29: 476. (1939)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium andesicola

cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur

Chiricahua Mountain larkspur

Stems

(33-)50-150(-280) cm;

base reddish, ± puberulent.

60-200 +cm;

base reddish or not, glabrous, glaucous.

Leaves

blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).

blade cordate to semicircular, 5-8 × 5-12 cm, nearly glabrous; ultimate lobes 5-16, width 3-20 mm, tips gradually tapered to mucronate apex;

midcauline leaf lobes more than 3 times longer than wide.

Inflorescences

10-40(-80)-flowered, open, narrowly pyramidal;

pedicel spreading, (1-)2-5 cm, ± puberulent;

bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.

20-80-flowered;

pedicel 1-2(-3) cm, puberulent;

bracteoles 1-3 mm from flowers, green to brown, linear-lanceolate, 3-6 mm, puberulent.

Flowers

sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm;

lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm;

hairs centered at base of cleft, short, sparse, yellow.

sepals purple, puberulent, lateral sepals spreading, 9-12 × 5-7 mm, spurs ascending ca. 45°, curved downward apically, purple, 10-13 mm, blunt tipped;

lower petal blades ± covering stamens, 4-6 mm, clefts 1.5-2.5 mm;

hairs centered, densest on inner lobes near base of cleft, white.

Fruits

erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.

12-15 mm, 3.5-4 times longer than wide, sparsely puberulent.

Seeds

unwinged;

seed coat cells with margins undulate, surfaces roughened.

unwinged;

seed coat cells elongate, surfaces pustulate.

= 16.

Delphinium andesicola

Phenology

Flowering spring–early summer.

Flowering summer–early fall.

Habitat

Slopes (often unstable) in chaparral

Meadows and coniferous woods

Elevation

50-1500 m (200-4900 ft)

2200-3200 m (7200-10500 ft)

Distribution

CA; Mexico (Baja California, Baja California Sur)

[WildflowerSearch map]

[BONAP county map]

Discussion

Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild.

Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study.

The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium andesicola, the westernmost representative of the southern Cordilleran complex, is found in the Chiricahua, Huachuca, Graham, and White mountains. Hybrids with Delphinium scopulorum are known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Exaltata

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. andesicola subsp. amplum

Name authority

Hooker: Bot. Mag., plate 4887. (1855)

Ewan: J. Washington Acad. Sci. 29: 476. (1939)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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