FNA: Delphinium barbeyi vs. Delphinium cardinale

	Delphinium barbeyi	Delphinium cardinale
	Barbey larkspur, Barbey's larkspur, subalpine larkspur, tall larkspur	cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur
Stems	50-150 cm; base green, glabrous.	(33-)50-150(-280) cm; base reddish, ± puberulent.
Leaves	blade round to reniform, 4-8 × 7-15 cm, glabrous; ultimate lobes 5-9, width 8-50 mm.	blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).
Inflorescences	10-50-flowered; pedicel 0.5-6 cm, glandular-puberulent; bracteoles 1-4(-8) mm from flowers, blue to green, awl-shaped, 5-14 mm, puberulent.	10-40(-80)-flowered, open, narrowly pyramidal; pedicel spreading, (1-)2-5 cm, ± puberulent; bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.
Flowers	sepals dark bluish purple, sparsely puberulent, lateral sepals forward pointing, 13-23 × 5-8 mm, spurs ascending ca. 45°, downcurved apically, 10-18 mm; lower petal blades ± covering stamens, 4-7 mm, clefts 2-3 mm; hairs centered, mostly near base of cleft, sparse elsewhere, white or yellow.	sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm; lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm; hairs centered at base of cleft, short, sparse, yellow.
Fruits	17-22 mm, 2.5-3 times longer than wide, puberulent.	erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.
Seeds	wing-margined; seed coat cells narrow but short, surfaces pustulate.	unwinged; seed coat cells with margins undulate, surfaces roughened.
2n	= 16.	= 16.

	Delphinium barbeyi	Delphinium cardinale
Phenology	Flowering summer.	Flowering spring–early summer.
Habitat	Subalpine and alpine sites in wet soils	Slopes (often unstable) in chaparral
Elevation	2500-4100 m (8200-13500 ft)	50-1500 m (200-4900 ft)
Distribution	AZ; CO; NM; UT; WY [WildflowerSearch map] [BONAP county map]	CA; Mexico (Baja California, Baja California Sur) [WildflowerSearch map] [BONAP county map]
Discussion	Delphinium barbeyi hybridizes extensively with D. glaucum in western Colorado and eastern Utah, where plants appearing to be hybrid [D. ×occidentale (S. Watson) S. Watson] are often far more common than plants of either putative parent. Several other names have been used for these plants, including D. elatum var. occidentale S. Watson, D. abietorum Tidestrom, and D. scopulorum subsp. occidentale (S. Watson) Abrams. Delphinium barbeyi is also known to hybridize with D. ramosum and D. sapellonis. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild. Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study. The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Exaltata	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms	D. exaltatum var. (e) barbeyi, D. occidentale var. barbeyi
Name authority	(Huth) Huth: Bull. Herb. Boissier 1: 335. (1893)	Hooker: Bot. Mag., plate 4887. (1855)
Web links	Local Web sites: SW CO Wildflowers WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium cardinale

Barbey larkspur, Barbey's larkspur, subalpine larkspur, tall larkspur

cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur

Stems

50-150 cm;

base green, glabrous.

(33-)50-150(-280) cm;

base reddish, ± puberulent.

Leaves

blade round to reniform, 4-8 × 7-15 cm, glabrous; ultimate lobes 5-9, width 8-50 mm.

blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline).

Inflorescences

10-50-flowered;

pedicel 0.5-6 cm, glandular-puberulent;

bracteoles 1-4(-8) mm from flowers, blue to green, awl-shaped, 5-14 mm, puberulent.

10-40(-80)-flowered, open, narrowly pyramidal;

pedicel spreading, (1-)2-5 cm, ± puberulent;

bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent.

Flowers

sepals dark bluish purple, sparsely puberulent, lateral sepals forward pointing, 13-23 × 5-8 mm, spurs ascending ca. 45°, downcurved apically, 10-18 mm;

lower petal blades ± covering stamens, 4-7 mm, clefts 2-3 mm;

hairs centered, mostly near base of cleft, sparse elsewhere, white or yellow.

sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm;

lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm;

hairs centered at base of cleft, short, sparse, yellow.

Fruits

17-22 mm, 2.5-3 times longer than wide, puberulent.

erect, 9-18 mm, 2.5-4 times longer than wide, glabrous.

Seeds

wing-margined;

seed coat cells narrow but short, surfaces pustulate.

unwinged;

seed coat cells with margins undulate, surfaces roughened.

= 16.

Delphinium cardinale

Phenology

Flowering summer.

Flowering spring–early summer.

Habitat

Subalpine and alpine sites in wet soils

Slopes (often unstable) in chaparral

Elevation

2500-4100 m (8200-13500 ft)

50-1500 m (200-4900 ft)

Distribution

AZ; CO; NM; UT; WY

[WildflowerSearch map]

[BONAP county map]

CA; Mexico (Baja California, Baja California Sur)

[WildflowerSearch map]

[BONAP county map]

Discussion

Delphinium barbeyi hybridizes extensively with D. glaucum in western Colorado and eastern Utah, where plants appearing to be hybrid [D. ×occidentale (S. Watson) S. Watson] are often far more common than plants of either putative parent. Several other names have been used for these plants, including D. elatum var. occidentale S. Watson, D. abietorum Tidestrom, and D. scopulorum subsp. occidentale (S. Watson) Abrams. Delphinium barbeyi is also known to hybridize with D. ramosum and D. sapellonis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild.

Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study.

The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Exaltata

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. exaltatum var. (e) barbeyi, D. occidentale var. barbeyi

Name authority

(Huth) Huth: Bull. Herb. Boissier 1: 335. (1893)

Hooker: Bot. Mag., plate 4887. (1855)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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