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Kern tarweed

tarplant, tarweed

Habit Annuals, 9–100 cm. Annuals, subshrubs, or shrubs, 4–120(–150) cm.
Stems

± solid or fistulose.

usually ± erect, rarely decumbent.

Leaves

proximal blades pinnatifid to toothed, faces ± hirsute and sometimes sparsely stipitate-glandular as well.

mostly cauline (annuals forming winter–spring rosettes, usually withering by flowering);

proximal opposite, most alternate;

sessile;

blades oblanceolate to linear or lance-linear, usually pinnatifid or toothed, sometimes serrate or entire, faces often hirsute or hispid-hirsute, sometimes villous, pilose, pubescent, canescent, strigose, or scabrous (often sessile- or stipitate-glandular as well) or glabrous (with scabrous or hispid margins).

Bracts

subtending heads sometimes overlapping proximal 0–1/2 of each involucre.

Involucres

± obconic, campanulate, hemispheric, or urceolate, 2–13+ mm diam.

Receptacles

flat to convex, glabrous or setulose, paleate (paleae falling, in 1 series between rays and disc in annuals, in 2–3+ series or subtending all or most disc florets in subshrubs and shrubs, connate or distinct, phyllary-like, more scarious).

Ray florets

(7–)8–12;

laminae pale yellow, 6–12 mm.

3–35, pistillate, fertile;

corollas deep or pale yellow.

Disc florets

10–21, all or mostly functionally staminate;

anthers yellow or brownish.

3–70, usually functionally staminate, seldom bisexual and fertile;

corollas yellow, tubes shorter than or about equaling funnelform throats, lobes 5, deltate (anthers usually reddish to dark purple or yellow, rarely maroon; styles glabrous proximal to branches).

Phyllaries

± evenly and minutely stipitate-glandular, including margins and apices, with non-glandular, non-pustule-based hairs as well.

falling, 3–35 in 1 series (± lanceolate to lance-attenuate or oblanceolate, herbaceous, each usually 1/2 enveloping subtended ray floret proximally, abaxially ± hirsute and sessile- or stipitate-glandular).

Heads

in crowded to open, corymbiform or paniculiform arrays.

radiate, usually in corymbiform or ± paniculiform arrays, sometimes in racemiform arrays or in glomerules.

Disc cypselae

usually 0;

pappi (of disc florets) usually of 1–15 elliptic, lance-linear, lanceolate, linear, oblong, quadrate, setiform, or subulate, entire, erose, fimbriate, fringed, or laciniate scales, sometimes 0 or coroniform (crowns of ± linear, sometimes fimbriate scales).

Ray cypselae

slightly obcompressed (adaxial sides flatter than abaxials), clavate (abaxially gibbous, often ± arcuate, basal attachments oblique, apices ± beaked, beaks offset adaxially, ascending, faces glabrous);

pappi 0.

Paleae

in 1 series.

Pappi

usually of 4–9 linear or oblong scales 0.8–1.1 mm, or of 1–5 subulate to setiform scales 0.1–0.9 mm, rarely 0.

Peduncular

bracts: pit-glands, tack-glands, and spines 0.

x

= 12 or 13.

2n

= 18.

Deinandra pallida

Deinandra

Phenology Flowering Mar–Jul.
Habitat Grasslands, open woodlands and shrublands, barrens, disturbed sites, sandy, silty, or clayey soils, often ± alkaline
Elevation 70–900 m (200–3000 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from USDA
w United States; nw Mexico
[BONAP county map]
Discussion

Deinandra pallida occurs in the southern San Joaquin Valley and the bordering Inner South Coast Ranges and southern Sierra Nevada foothills.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 21 (16 in the flora).

Deinandra is treated here as distinct from Hemizonia; Deinandra is more closely related to Calycadenia, Centromadia, Holocarpha, and Osmadenia than to Hemizonia in the strict sense (S. Carlquist et al. 2003). As circumscribed here (following B. G. Baldwin 1999b), Deinandra comprises all taxa included in Hemizonia sect. Madiomeris by B. D. Tanowitz (1982) plus the shrubs and subshrubs constituting the informal “Fruticosae” or “Zonamra” (J. Clausen 1951; D. D. Keck 1959b). All are self-incompatible except D. arida and D. mohavensis. Five species (D. frutescens, D. greeneana, D. martirensis, D. palmeri, D. streetsii), none annuals, are known only from Baja California, Mexico. Most species north of Mexico are cross-incompatible or ± intersterile (J. Clausen 1951).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Subshrubs or shrubs; paleae in 2 series or in 3+ series or throughout receptacles
→ 2
1. Annuals; paleae in 1 series (between ray and disc florets)
→ 3
2. Paleae in 2 series (1 between ray and disc florets, 1 between outermost and adjacent discflorets); ray florets (11–)13(–20); anthers reddish to dark purple
D. clementina
2. Paleae in 3+ series or throughout receptacles; ray florets (4–)8; anthers yellow or brownish
D. minthornii
3. Ray florets 3–5; disc florets 3–6; pappi of 5–12 scales
→ 4
3. Ray florets (5–)8–35; disc florets 8–70; pappi 0, or of 1–14 scales, or coroniform (reputedly sometimes rudimentary)
→ 8
4. Ray florets 3(–4); disc florets 3(–4)
D. lobbii
4. Ray florets 5; disc florets 6
→ 5
5. Phyllaries usually sessile-glandular, at least near margins, rarely stipitate-glandular (stalks shorter than glands); anthers reddish to dark purple
→ 6
5. Phyllaries stipitate-glandular (stalks often equal to or longer than glands); anthers yellow or brownish (reddish to dark purple or maroon in some, mostly s Californian, D. kelloggii)
→ 7
6. Bracts subtending heads usually overlapping at least proximal 1/2 of each involucre; phyllaries glandular near margins, sometimes with non-glandular, non-pustule-based hairs as well
D. fasciculata
6. Bracts subtending heads usually overlapping proximal 0–1/2 of each involucre; phyllaries ± evenly glandular and with pustule-based hairs, atleast on midribs
D. pentactis
7. Proximal leaves usually pinnatifid to toothed (rarely entire); heads in open, paniculiform arrays; pappi of linear to oblong, entire or fringed scales
D. kelloggii
7. Proximal leaves usually entire, sometimes serrate; heads usually in glom-erules; pappi of irregular, erose scales
D. mohavensis
8. Anthers yellow or brownish
→ 9
8. Anthers reddish to dark purple
→ 12
9. Faces of proximal leaves glabrous or hispid-hirsute and stipitate-glandular; disc florets 17–60; pappi usually 0, rarely of 1–5 linear to setiform scales 0.1–0.6 mm or, reputedly, rudimentary
→ 10
9. Faces of proximal leaves ± hirsute and stipitate-glandular; disc florets 10–21; pappi usually of 4–13 scales, sometimes coroniform (in D. bacigalupii), rarely 0 (in Deinandra pallida)
→ 11
10. Stems solid; leaves (proximal) hispid-hirsute and stipitate-glandular; disc florets 17–25
D. arida
10. Stems fistulose; leaves (proximal) glabrous (margins and midribs sometimes scabrous or hispid); disc florets 28–60
D. halliana
11. Ray laminae pale yellow, 6–12 mm
D. pallida
11. Ray laminae deep yellow, 2–4 mm
D. bacigalupii
12. Disc florets all or mostly bisexual
D. floribunda
12. Disc florets all or mostly functionally staminate
→ 13
13. Ray florets 15–35; pappi 0, or coroniform (irregular crowns of entire, erose, orlaciniate scales 0.1–0.9 mm)
D. corymbosa
13. Ray florets (7–)8–13; pappi of 4–14 elliptic, lanceolate, linear, oblong, or quadrate, barely fringed to erose scales 0.5–2 mm
→ 14
14. Phyllaries sessile- and stipitate-glandular, mostly near proximal margins and on apices
D. conjugens
14. Phyllaries ± evenly stipitate-glandular, including margins and apices
→ 15
15. Ray florets (7–)8(–10); disc florets 8–14(–15) (basal leaves sometimes present at flowering in South Coast Ranges)
D. paniculata
15. Ray florets 8–13(–15); disc florets 11–32
D. increscens
Source FNA vol. 21, p. 286. FNA vol. 21, p. 280. Authors: Bruce G. Baldwin, John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Madiinae > Deinandra Asteraceae > tribe Heliantheae > subtribe Madiinae
Sibling taxa
D. arida, D. bacigalupii, D. clementina, D. conjugens, D. corymbosa, D. fasciculata, D. floribunda, D. halliana, D. increscens, D. kelloggii, D. lobbii, D. minthornii, D. mohavensis, D. paniculata, D. pentactis
Subordinate taxa
D. arida, D. bacigalupii, D. clementina, D. conjugens, D. corymbosa, D. fasciculata, D. floribunda, D. halliana, D. increscens, D. kelloggii, D. lobbii, D. minthornii, D. mohavensis, D. pallida, D. paniculata, D. pentactis
Synonyms Hemizonia pallida
Name authority (D. D. Keck) B. G. Baldwin: Novon 9: 469. (1999) Greene: Fl. Francisc. 4: 424. (1897)
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