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cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

and leaves with non-Kranz anatomy (C3).

Leaves

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

spikes;

rachilla persistent, wingless.

Spikelets

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads;

floral scales deciduous, 2-keeled or folded (conduplicate).

Flowers

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

stamens 1–3;

stigmas 3.

Achenes

biconvex, flattened, or trigonous.

trigonous, plano-convex, or terete.

Cyperus

Cyperus subg. Pycnostachys

Distribution
map from USDA
Pantemperate and tropical
[BONAP county map]
Worldwide
Discussion

Species ca. 600 (96 in the flora).

Etymology: Greek kupeiros, name for Eurasian Cyperus longus Linnaeus

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 150 (22 in the flora).

The greatest species diversity of Cyperus subg. Pycnostachys is found in the lowland forests of Central America, South America, Africa, and Malesia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Cyperaceae Cyperaceae > Cyperus
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Synonyms C. subg. Protocyperus
Name authority Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754) C. B. Clarke: in J. D. Hooker, Fl. Brit. India 6: 597. (1893)
Source FNA vol. 23, p. 141. Treatment authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *. FNA vol. 23. Treatment authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
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