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green flatsedge

smooth cyperus, smooth flatsedge, smooth nutgrass

Habit Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. Herbs, perennial, rhizomatous.
Culms

trigonous, (20–)60–70(–100) cm × 2.4–6.9 mm, scabrid on angles.

single or close together, trigonous to ± terete, (1–)8–30(–60) cm × (0.3–)0.6–1.5(–2.2) mm, glabrous.

Leaves

4–12, flat to V-shaped, with readily visible cross ribs especially on abaxial surface, 20–50 cm × 3–14 mm.

blades present or reduced to sheaths, base of culm covered with reddish sheath 5–25 mm, distally short toothed 1–2 mm, or leaf blades involute, 1–7 cm × 0.3–1(–1.6) mm.

Inflorescences

heads digitate to hemispheric, 1–3.5 cm diam.;

rays 6–12, 1–9(–14) cm; 2d order rays often present, (0.5–)1–3 cm; 3d order rays occasionally present, 5–12 mm;

bracts 4–8, horizontal to ascending at 30°, with prominent cross ribs,V-shaped, (1.5–)3–50(–75) cm × 0.5–13 mm; 2d order bracts 3–10 × 1–2 mm.

heads appearing lateral, spikes ± digitate, 5–15 × 5–15 mm;

rays absent, sometimes 1, to 2 mm;

bracts (1–)2(–3), longer bracts erect, appearing as continuation of culm, shorter bract(s) horizontal or ascending, longer bracts (1–)2–8(–12) cm × 0.5–3 mm, shorter (0.3–)1–4.5 cm × 0.2–1 mm.

Spikelets

10–40(–50), oblong to linear-lanceoloid, 5–18 × 2–3.2 mm;

floral scales 10–40, pale grayish brown, or greenish, proximally greenish or brownish, oblanceolate, 2-keeled, weakly to distinctly 1-ribbed, proximal half 2-ribbed, (1.3–)1.5–2(–2.4) × 0.9–1.4 mm, apex acute (sometimes mucronulate), apically glabrous, occasionally scabridulous.

(1–)2–7(–14), whitish green to reddish, compressed-turgid, ellipsoid to oblong-lanceoloid, 4–7(–12) × (1.6–)2–3(–3.6) mm;

floral scales (8–)12–22, off-white, whitish with red speckles, or with single red spot 0.5 mm wide, medially whitish, oblong-laterally ribless or weakly 2-ribbed, medially weakly 1–3-ribbed, obovate to orbiculate, 1.5–2 × (1.3–)1.6–2 mm.

Flowers

stamens 1–2;

anthers 0.8–1.4 mm;

styles 0.8–1 mm;

stigmas 0.6–0.8 mm.

anthers (0.5–)0.7–1 mm, connective apex reddish, 0.1–0.2 mm;

styles 0.8–1.2 mm;

stigmas 0.7–1 mm.

Achenes

brown, slightly to distinctly stipitate, ellipsoid (2.7–4.1 times as long as wide), (0.9–)1.2–1.5 × 0.3–0.4 mm, apical beak 0.1–0.5 mm, surfaces glabrous.

gray, glossy, oblong-ellipsoid to ovoid, (1.2–)1.3–1.5(–1.8) × (0.7–)0.8–1(–1.2) mm, base cuneate to stipelike, apex apiculate, surfaces finely reticulate to papillate.

Cyperus virens

Cyperus laevigatus

Phenology Fruiting spring–fall. Fruiting summer.
Habitat Wet pastures, marshes, roadside ditches Alkaline grasslands, hotsprings, ditches, brackish marshes
Elevation 0–1000 m (0–3300 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; Mexico; Central America; South America
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[BONAP county map]
from FNA
AZ; CA; NC; TX; Mexico; South America; West Indies; Europe; Asia; Africa
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Discussion

Cyperus virens was found as waif in California in the 1800s (M. F. Denton 1978b; G. C. Tucker 1993b).

Cyperus virens is readily distinguished from other species of subg. Pycnostachys by its trigonous culms, scabrid angles, and leaves and inflorescence bracts conspicuously septate by numerous cross ribs between the main ribs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus laevigatus is one of only two species of subg. Juncellus in the New World. It can be distinguished from all other New World species of Cyperus by its two stigmas and disclike, strongly flattened achene, which is borne with its face toward the rachilla. Species of subg. Pycreus, especially C. niger, have been misidentified as C. laevigatus, probably because of the dark, shiny spikelets. All species of subg. Pycreus have biconvex achenes like C. laevigatus; in species of subg. Pycreus the achene is attached with an edge toward the rachilla; in C. laevigatus the achene faces the rachilla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 153. FNA vol. 23, p. 164.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Juncellus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms Acorellus laevigatus, Chlorocyperus laevigatus, Juncellus laevigatus, Pycreus laevigatus
Name authority Michaux: Fl. Bor.-Amer. 1: 28. (1803) Linnaeus: Mant. Pl. 2: 179. (1771)
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