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green flatsedge

woodrush flatsedge

Habit Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm.
Culms

trigonous, (20–)60–70(–100) cm × 2.4–6.9 mm, scabrid on angles.

1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm.

Leaves

4–12, flat to V-shaped, with readily visible cross ribs especially on abaxial surface, 20–50 cm × 3–14 mm.

(1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm.

Inflorescences

heads digitate to hemispheric, 1–3.5 cm diam.;

rays 6–12, 1–9(–14) cm; 2d order rays often present, (0.5–)1–3 cm; 3d order rays occasionally present, 5–12 mm;

bracts 4–8, horizontal to ascending at 30°, with prominent cross ribs,V-shaped, (1.5–)3–50(–75) cm × 0.5–13 mm; 2d order bracts 3–10 × 1–2 mm.

heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.;

rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm;

bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm.

Spikelets

10–40(–50), oblong to linear-lanceoloid, 5–18 × 2–3.2 mm;

floral scales 10–40, pale grayish brown, or greenish, proximally greenish or brownish, oblanceolate, 2-keeled, weakly to distinctly 1-ribbed, proximal half 2-ribbed, (1.3–)1.5–2(–2.4) × 0.9–1.4 mm, apex acute (sometimes mucronulate), apically glabrous, occasionally scabridulous.

(1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm;

floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous.

Flowers

stamens 1–2;

anthers 0.8–1.4 mm;

styles 0.8–1 mm;

stigmas 0.6–0.8 mm.

stamen 1;

anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm;

styles 0.2–0.4 mm;

stigmas 0.4–0.6 mm.

Achenes

brown, slightly to distinctly stipitate, ellipsoid (2.7–4.1 times as long as wide), (0.9–)1.2–1.5 × 0.3–0.4 mm, apical beak 0.1–0.5 mm, surfaces glabrous.

brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate.

Cyperus virens

Cyperus entrerianus

Phenology Fruiting spring–fall. Fruiting summer.
Habitat Wet pastures, marshes, roadside ditches Roadside ditches, marshes
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus virens was found as waif in California in the 1800s (M. F. Denton 1978b; G. C. Tucker 1993b).

Cyperus virens is readily distinguished from other species of subg. Pycnostachys by its trigonous culms, scabrid angles, and leaves and inflorescence bracts conspicuously septate by numerous cross ribs between the main ribs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America.

Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 153. FNA vol. 23.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Pycnostachys
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. luzulae var. entrerianus
Name authority Michaux: Fl. Bor.-Amer. 1: 28. (1803) Boeckeler: Flora 61: 139. (1878)
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