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tropical flatsedge

cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, annual or short-lived perennial, annuals cespitose; rhizomes absent. Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

2–6, trigonous, (10–)35–80 cm × 0.5–2(–4) mm, scabridulous with retrorse prickles (infrequently glabrous or rarely also with extrorse prickles).

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

Leaves

3–9, V-shaped, (12–)25–45(–65) cm × 2–10 mm;

blades and bracts without cross ribs.

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

heads umbellate, 10–20(–30) mm diam.;

rays 4–12, 1–6 cm, minutely scabridulous with retrorse prickles; 2d order rays often present, 1–2(–3) cm; 3d order rays rarely present, 0.5–1.5 cm;

bracts 3–8, approximately horizontal to ascending at 30°, V-shaped to flat, 2–15(–34) cm × 1–4(–5) mm.

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

Spikelets

(6–)15–40(–65), greenish white, linear to linear-oblong, (3–)4–12(–15) × 1.5–2.5 mm;

floral scales 10–50(–65), laterally pale yellow, light brown, or reddish brown, 2-keeled, medially 3-ribbed, lanceolate, 1–1.5 × 0.8–0.9 mm, distinctly reticulate, often scabridulous near apex.

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

Flowers

stamen 1;

anthers 0.5 mm;

styles 0.8–1.1 mm;

stigmas 0.5 mm.

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

Achenes

brown to reddish brown, slightly stipitate, narrowly ellipsoid, 0.7–0.9 × 0.2–0.4 mm, apex apiculate, surfaces papillate or obscurely reticulate to rugulose.

biconvex, flattened, or trigonous.

Cyperus surinamensis

Cyperus

Phenology Fruiting late spring–early fall.
Habitat Moist sunny areas with disturbed soils
Elevation 0–200 m (0–700 ft)
Distribution
from FNA
AL; AR; FL; GA; KS; LA; MS; OK; SC; TN; TX; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
from USDA
Pantemperate and tropical
[BONAP county map]
Discussion

Cyperus surinamensis is distinguished readily from all other species of the genus in the New World by the usual presence of retrorse prickles on the culms and rays.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 600 (96 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Source FNA vol. 23, p. 153. FNA vol. 23, p. 141. Authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Name authority Rottbøll: Descr. Pl. Rar., 20. (1772) Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754)
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