Cyperus schweinitzii |
Cyperus niger |
|
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sand flatsedge, Schweinitz's flatsedge |
black flatsedge, brown cyperus |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, rhizomatous. |
Culms | trigonous, (10–)20–50 cm × (0.6–)1–2 mm, glabrous or more often scabridulous, either in distal 1/2 or over entire length; basal sheaths, 1–3, light brown, 1–4 cm, papery. |
solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
Leaves | flat to V-shaped, (2–)20–35 cm × 2–6 mm, margins and keel minutely scabridulous; proximal leaves bladeless. |
(2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
Inflorescences | spikes ± umbellate to slenderly (infrequently broadly) ovoid, 10–25 × 7–16 mm; rays 3–5, 1.5–8(–15) cm; bracts 3–7, erect (or declined at most 25–30° from vertical), (2.5–)8–20(–25) cm × (0.5–)1.5–6 mm; rachilla ± deciduous, wingless. |
spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
Spikelets | (1–)5–10(–20), oblong to linear-oblong, quadrangular, 7–10(–18) × (2.8–)3.2–4(–4.5) mm; floral scales deciduous, (1–)5–10(–14), spreading, laterally stramineous or dull whitish, sometimes also red-spotted, medially green, laterally 2–3-ribbed, medially 3-ribbed, broadly ovate to oblong-ovate, (2.3–)2.8–3.2 × (2–)2.4–3.2 mm, apex broadly rounded, often clear-erose, cusp 0.1–0.4 mm in proximal scales, 0.3–1 mm in distal scales. |
(3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
Flowers | anthers 0.8–1.4 mm; styles 0.1–0.3 mm; stigmas 1.5–3 mm. |
stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
Achenes | light to dark brown or nearly black, broadly ellipsoid, infrequently obovoid, 2–2.4 × 0.9–1.4 mm, base cuneate to ± stipelike, apex obtuse to broadly rounded, surfaces papillose. |
brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
2n | = 166. |
|
Cyperus schweinitzii |
Cyperus niger |
|
Phenology | Fruiting late spring–early summer. | Fruiting summer. |
Habitat | Riverbanks, sand bars, lakeshores, sand dunes, sandy openings in woods | Swamps, ditches, wet pastures, seeps in montane forests |
Elevation | 0–1000 m (0–3300 ft) | 0–1200 m (0–3900 ft) |
Distribution |
CO; IA; IL; KY; MA; MI; MN; MO; MT; NJ; NM; NY; OH; OK; PA; SD; TX; UT; WA; WI; WY; AB; MB; ON; QC; SK; Mexico (Coahuila)
|
AZ; CA; NM; OK; TX; Mexico; Central America; South America
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Discussion | Cyperus schweinitzii is introduced, but not naturalized, in Massachusetts and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 174. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. alterniflorus | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus |
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York 3: 276. (1836) | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) |
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