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chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge

false flatsedge, false nutsedge, straw color cyperus, straw-color flatsedge, straw-colour flatsedge

Habit Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. Herbs, perennial (often flowering first year), rhizomatous.
Culms

trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous.

basally swollen and cormlike, trigonous, (1–)20–40(–90) cm × 1–6 mm, glabrous or few prickles subapically.

Leaves

V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm.

flat, (10–)20–40 cm × 1–4(–8) mm, margins and midribs glabrous or scabridulous.

Inflorescences

spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide.

spikes 1–4, loosely ovoid to oblong-cylindric; (6–)10–28(–50) × 10–40 mm;

rays sometimes absent, usually (1–)3–6(–8), 0.5–7(–25) cm;

if absent, inflorescence a congested head; 2d order rays sometimes present, 1–4(–9) cm;

rachis 8–18(–25) mm;

bracts (3–)5–7(–10), ascending at 30–45°, cross ribs sometimes conspicuous, (1–)10–30(–45) cm × (1–)2–4(–12) mm;

rachilla ± deciduous, wings 0.4–0.5 mm wide.

Spikelets

(2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse.

(5–)12–50, linear-lanceoloid, compressed, 5–30 × 0.6–0.9 mm;

floral scales ± deciduous, 3–11, appressed, medially green, laterally stramineous to pale brown, 3–4-ribbed laterally, (1–)3-ribbed medially, oblong-obovate, 3.2–4.5(–6) × 1–1.8 mm, apex ± acute, overlapping lower 1/4–1/2 of next scale.

Flowers

anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm.

anthers 0.3–0.5 mm;

styles 1–2 mm;

stigmas 3–4 mm.

Achenes

black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate.

purplish brown, ± stipitate, narrowly oblong, (1.5–)1.8–2.4 × 0.5–0.6 mm, apex apiculate, acute, surfaces puncticulate.

Cyperus rotundus

Cyperus strigosus

Phenology Fruiting summer–fall. Fruiting summer.
Habitat Croplands, disturbed soils usually Pond shores, ditches, damp, disturbed soils, sometimes weeds of gardens or croplands
Elevation 0–400 m (0–1300 ft) 0–1500 m (0–4900 ft)
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; MB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations.

Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus.

Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus strigosus is usually among the more common Cyperus species throughout its range. Small individuals flowering the first year may be difficult to distinguish from C. polystachyos, C. odoratus, C. erythrorhizos, and C. esculentus, which may be sympatric. Cyperus strigosus has trigonous achenes and three stigmas, unlike C. polystachyos; C. strigosus has flattened spikelets, unlike the subcylindric ones of C. odoratus; C. strigosus has floral scales usually 3 mm or more, unlike the smaller (1.2–1.5 mm) ones of C. erythrorhizos; C. strigosus has deciduous floral scales and a cormlike stem base with stolons, unlike C. esculentus.

Cyperus strigosus appears to be closely related to the neotropical C. camphoratus Liebmann; both species have deciduous floral scales and deciduous spikelets (G. C. Tucker 1994). Plants segregated as C. stenolepis cannot be distinguished consistently from C. strigosus on any single charcter, rather they appear to be merely large individuals of C. strigosus with long floral scales and frequently septate inflorescence bracts (M. L. Horvat 1941).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 169. FNA vol. 23, p. 184.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. hansenii, C. stenolepis, C. strigosus var. hansenii, C. strigosus var. stenolepis, Mariscus stenolepis, Mariscus strigosus
Name authority Linnaeus: Sp. Pl. 1: 45. (1753) Linnaeus: Sp. Pl. 1: 47. (1753)
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