Cyperus rotundus |
Cyperus plukenetii |
|
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chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
Plukenet's flatsedge, Plukenet's sedge |
|
Habit | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. | Herbs, perennial, cespitose. |
Culms | trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
basally cormlike, obtusely trigonous to subterete, 25–100 cm × 2–4 mm, densely minutely scabridulous. |
Leaves | V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
inversely W-shaped, 20–70 cm × 4–7 mm, adaxial surface densely hirtellous. |
Inflorescences | spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
spikes turbinate, 10–12 × 6–9 mm, ± as long as wide; rays 6–12, 1–30 cm (longest bract equaling or shorter than longest ray), harshly scabrid; bracts 3–10, horizontal to ascending at 30°, inversely W-shaped, 6–27 cm × 3–6 mm; rachilla persistent, wings hyaline, 0.5 mm wide, covering 1/2 of mature achene. |
Spikelets | (2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
30–60, proximal spikelets reflexed nearly parallel to ray, distal ones spreading, linear, ± terete, 6–8 × 1–1.4 mm; floral scales persistent, 1(–2), appressed, stramineous, often suffused with reddish purple, laterally 4–6-ribbed, ovate-lanceolate, 4–4.5 × 1.4–1.6 mm, apex straight, excurved mucronate or cuspidate; terminal scale spinose. |
Flowers | anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
anthers 0.5–1 mm; styles 1.5–2 mm; stigmas 2 mm. |
Achenes | black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
dark brown, narrowly oblong, (2.4–)2.7–2.9(–3.2) × 0.7–0.8 mm, base cuneate, apex obtuse, apiculate, surfaces puncticulate. |
Cyperus rotundus |
Cyperus plukenetii |
|
Phenology | Fruiting summer–fall. | Fruiting summer (Jul–Aug). |
Habitat | Croplands, disturbed soils usually | Dry sand hills, xeric upland woods |
Elevation | 0–400 m (0–1300 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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AL; AR; DE; FL; GA; KY; LA; MD; MO; MS; NC; NJ; OK; SC; TN; TX; VA
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Discussion | Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus plukenetii is distinguished by its tight, burlike spikes, few-flowered spikelets, and densely pubescent culms, leaves, and rays. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 169. | FNA vol. 23, p. 180. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Fernald: Rhodora 47: 110, plate 877. (1945) |
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