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chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge
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plains flatsedge
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| Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. |
Herbs, perennial, cespitose, rhizomatous; base cormlike; rhizomes scaly, 12 cm × 2 mm. |
trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
trigonous, 20–55 cm × 1.2–11.8 mm, glabrous. |
V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
V-shaped, 12–30 cm × 2–3 mm. |
spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
heads digitate, 12–20 mm diam.; rays 7–12, 1–13 cm; 2d order rays 1–3 cm (sometimes absent); bracts 3–5, longest ± erect, V-shaped, 5–12(–18) cm × 2–3 mm. |
(2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
(8–)20–30, linear-lanceoloid, compressed, 5–10(–14) × 1.4–1.7 mm; floral scales (8–)16–26, laterally brown to reddish brown, medially green, laterally 1-ribbed, medially 3-ribbed, broadly elliptic, 2–2.5 × 1.2–1.6 mm, apex with slightly excurved cusp 0.2–0.3 mm. |
anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
stamens 3; anthers 1 mm, connective apex reddish, subulate, 0.1 mm; styles 1–1.5 mm; stigmas 1.5 mm. |
black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
white to light brown, sessile, ellipsoid, 0.7–0.8 × 0.25–0.35 mm, apex obtuse, apiculate, surfaces puncticulate. |
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| Fruiting summer–fall. |
Fruiting early summer (May–Jun). |
| Croplands, disturbed soils usually |
Permanent pools and wet swales between sand dunes |
| 0–400 m (0–1300 ft) |
1200 m (3900 ft) |
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AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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TX |
Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Cyperus onerosus is apparently restricted to Andrews and Winkler counties in Texas. This interesting endemic is most similar to Cyperus dentatus; it lacks tubers and apparently is not proliferous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 23, p. 169. |
FNA vol. 23, p. 151. |
| Cyperaceae > Cyperus > subg. Cyperus |
Cyperaceae > Cyperus > subg. Pycnostachys |
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens |
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens |
| Linnaeus: Sp. Pl. 1: 45. (1753) |
M. C. Johnston: SouthW. Naturalist 9: 308. (1964) |
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CA
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