Cyperus rotundus |
Cyperus lentiginosus |
|
---|---|---|
chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
Latin American flatsedge |
|
Habit | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. | Herbs, perennial, cespitose, rhizomatous. |
Culms | trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
trigonous, (1–) 20–40(–75) cm × 1–1.5(–2.5) mm, glabrous. |
Leaves | V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
inversely W-shaped, 10–40(–70) cm × (1–)3–6(–10) mm. |
Inflorescences | spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
spikes 1(–3), broadly and loosely ovoid to oblong-ellipsoid, (15–)20–30(–50) × 20–30(–45) mm; bracts (3–)5–7(–10), ascending at 30(–45)°, inversely W-shaped, 10–30(–50) cm × (1–)3–5(–11) mm; rays (2–)3–6(–12), (1–)3–10(–17) cm; rachilla deciduous, wings ca. 0.3 mm wide. |
Spikelets | (2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
(10–)25–50(–100), ± terete, (appearing compressed due to excurved apices of floral scales), (8–)12–15(–24) × (0.8–)1.1–1.4(–1.8) mm; floral scales persistent, (4–)6–8(–14), golden brown, off-white to stramineous, densely red-glandular punctate, green medially, 3–4-ribbed laterally, 3–5-ribbed medially, oblong-lanceolate, (3.2–)4–4.5 × 1.2–1.6 mm, apex blunt, mucronulate to mucronate, mucro 0.2–0.5 mm; proximal scales mucronulate, distal scales mucronate; terminal scale conduplicate. |
Flowers | anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
anthers 0.5–0.8(–1.4) mm; styles 0.8–1 mm; stigmas 1–2 mm. |
Achenes | black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
brown, ± sessile to stipitate, oblong-ellipsoid, 1.7–2 × 0.6–0.7 mm, 0.1(–0.2) × 0.2 mm, apex rounded, apiculate from dark purple style base, surfaces papillose. |
Cyperus rotundus |
Cyperus lentiginosus |
|
Phenology | Fruiting summer–fall. | Fruiting spring–summer. |
Habitat | Croplands, disturbed soils usually | Thickets, open woods |
Elevation | 0–400 m (0–1300 ft) | 0–50 m (0–200 ft) |
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
|
FL; TX; Mexico; West Indies |
Discussion | Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus lentiginosus has been treated as a variety of C. tenuis (G. Kükenthal 1935–1936). The two taxa differ in the longer scales with cuspidate apices of C. lentiginosus, the inversely W-shaped leaves and bracts (those of C. tenuis are V-shaped), and the open spikes (those of C. tenuis are dense). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 169. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. strigosus var. gracilis, C. tenuis var. lentiginosus | |
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Millspaugh & Chase: Publ. Field Columbian Mus., Bot. Ser. 3: 74. (1903) |
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