Cyperus rotundus |
Cyperus involucratus |
|
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chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
umbrella flatsedge, umbrella-plant |
|
Habit | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. | Herbs, perennial, cespitose, rhizomatous. |
Culms | trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
trigonous, 30–150 cm × 1–5(–8) mm. |
Leaves | V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
bladeless. |
Inflorescences | spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
heads digitate, 15–30(–36) mm diam.; rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm; bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm. |
Spikelets | (2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm; floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute. |
Flowers | anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
stamens 3; anthers 0.7–1 mm; styles 0.5–1 mm; stigmas 0.6–1 mm. |
Achenes | black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate. |
Cyperus rotundus |
Cyperus involucratus |
|
Phenology | Fruiting summer–fall. | Fruiting early summer–fall. |
Habitat | Croplands, disturbed soils usually | Damp, disturbed soils, ditches, stream banks |
Elevation | 0–400 m (0–1300 ft) | 0–100(–800) m (0–300(–2600) ft) |
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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CA; FL; LA; TX; e Africa [Introduced in North America]
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Discussion | Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997). Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 169. | FNA vol. 23, p. 150. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Rottbøll: Descr. Pl. Rar., 22. (1772) |
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