Cyperus rotundus |
Cyperus fugax |
|
---|---|---|
chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
withering flatsedge |
|
Habit | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. | Herbs, annual, cespitose. |
Culms | trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
(1–)10–30(–60) together, roundly trigonous proximally, trigonous distally, (0.5–)3–6(–12) cm × (0.2–)0.4–0.8 mm, glabrous. |
Leaves | V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
V-shaped, (1–)3–5(–10) cm × (0.5–)1–2 mm. |
Inflorescences | spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
spikes loosely ovoid, 8–15(–25) mm wide; rachis 1–3 mm; rays (1–)2–4(–6), (0.5–)1–3(–6) cm; 2d order rays absent; bracts (1–)2–4(–5), ascending at 30–45°, V-shaped, (1–)2.5–6(–11) cm × 0.5–1.8 mm. |
Spikelets | (2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
(2–)4–6(–10), linear-lanceoloid, strongly flattened, (5–)8–12(–18) × 1.2–1.5 (1.8) mm; floral scales (5–)10–16(–32), laterally clear, stramineous or light reddish brown, medially green, laterally ribless, medially 3(–5)-ribbed, ovate, (1.2–)1.3–1.6(–2) × 1–1.2(–1.4) mm, apex obtuse, mucronate to minutely mucronulate. |
Flowers | anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
stamens 2; anthers 0.1–0.2 mm; styles 0.1–0.3(–0.4) mm; stigmas 0.3–0.4 mm. |
Achenes | black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
light to dark brown, glossy or iridescent, stipitate, oblong-obovoid, 0.9–1.1 × 0.4–0.5 mm, stipe 0.1 mm, apex truncate, apiculate, surfaces punctate. |
Cyperus rotundus |
Cyperus fugax |
|
Phenology | Fruiting summer–fall. | Fruiting early summer. |
Habitat | Croplands, disturbed soils usually | Wet, disturbed sandy soils in full sun |
Elevation | 0–400 m (0–1300 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
|
LA; TX; Central America; West Indies (Cuba); South America (Ecuador) |
Discussion | Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus fugax is known from Louisiana from an early nineteeth century collection from “western Louisiana” (Hale s.n., TCD). Cyperus fugax has been treated as a form of C. polystachyos (G. Kükenthal 1935–1936; A. B. Ayers 1946); nevertheless, specific status is appropriate. The most conspicuous differences are the annual habit and small size (less than 12 cm) of C. fugax; C. polystachyos is a perennial of moderate size (mostly taller than 20 cm). Other differences are summarized below. Cyperus polystachyos is characterized as a shortly rhizomatous perennial; floral scales oblong, 1.8–2.4 mm, closely imbricate; styles 0.6–1 mm; stigmas 1.4–2 mm; anthers 0.6–0.8 mm; achenes substipitate to cuneate, slightly compressed laterally. Cyperus fugax is characterized as a nonrhizomatous annual; floral scales ovate, 1.3–1.6 mm, loosely imbricate; styles 0.1–0.2 mm; stigmas 0.3–0.4 mm; filaments 1.2–1.4 mm; anthers 0.1–0.2 mm; achenes stipitate, strongly compressed laterally. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 169. | FNA vol. 23, p. 159. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. inconspicuus, C. tenellus | |
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Liebmann: Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 2: 196. (1850) |
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