Flora of North America species comparison

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Cyperus rotundus

Cyperus diandrus

chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge

umbrella flat sedge
Habit Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. Herbs, annual, cespitose, with fibrous roots.
Culms

trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous.

trigonous, 4–25(–35) cm, glabrous.
Leaves

V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm.

flat, 5–20 cm × 1.5–3 mm.
Inflorescences

spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide.

heads ± digitate, 12–22 mm diam.;

rays 3–6, 1–6 cm;

bracts 1–4, ± horizontal, flat, 1.5–15 cm × 1.5–3 mm.
Spikelets

(2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse.

6–12, oblong-lanceoloid, compressed, 5–10 × 2.8–3.2 mm;

floral scales 8–28, closely imbricate, stramineous to light brown or clear (marginal band bright red, spikelet thus appearing dark margined), laterally ribless, oblong to ovate, 2.5–3 × 1.6–1.9 mm.
Flowers

anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm.

stamens 2(–3);

anthers 0.3 mm, connective apex reddish, subulate, 0.1–0.2 mm;

styles 0.3–0.5 mm;

stigmas 2.2–3.1 mm.
Achenes

black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate.

brown, network of ridges forming isodiametric or square cells, sessile or stipitate, obovoid to ovoid, 1–1.2 × 0.5–0.6 mm, stipe, if present, 0.1 mm, apex obtuse, apiculate, surfaces minutely punctate.

Cyperus rotundus

Cyperus diandrus
Phenology Fruiting summer–fall. Fruiting summer.
Habitat Croplands, disturbed soils usually Emergent shorelines, in sandy, peaty, or slightly brackish areas (though seldom where disturbed)
Elevation 0–400 m (0–1300 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DE; IA; IL; IN; MA; MD; ME; MI; MN; MO; NE; NH; NJ; NY; PA; RI; VA; VT; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations.

Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus.

Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 23, p. 169. FNA vol. 23, p. 161.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae > Cyperus > subg. Pycreus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms Pycreus diander
Name authority Linnaeus: Sp. Pl. 1: 45. (1753) Torrey: Cat. Pl. New York, 90. (1819)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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