Cyperus rotundus
Cyperus dentoniae
chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge
Denton's flatsedge, hairy flatsedge
trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous.
1–4, trigonous, (2–)20–50(–80) cm × (0.8–)1.5–2(–2.8) mm, basally glabrous or nearly so, apically hispidulous, on faces and angles distally, especially so immediately proximal to bracts.
V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm.
2–5(–7), V-shaped, (5–)15–30(–45) cm × (1.5–)2.5–5(–8) mm.
spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide.
spike 1 (occasionally 1–2 smaller sessile spikes at base), densely cylindric, (7–)11–20(–25) × (5–) 7–10(–12) mm;
rays 3–6(–10), (1–)3–6(–18) cm, rays and rachis hispidulous;
bracts 4–8(–10), ascending at 30–60°, V-shaped, (2–)10–25(–45) cm × (1–)3–6(–8) mm;
rachilla deciduous, wings persistent, 0.3–0.4 mm wide.
(2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse.
(20–)70–120(–140), oblong-ellipsoid to oblong-lanceoloid, ± terete to quadrangular, (2.2–)3.4–4.5(–5.6) × (0.8–)0.9–1.2(–1.3) mm;
distal spikelet spreading or ascending;
floral scales 1(–3), appressed, stramineous, red-spotted, strongly 3(–4)-ribbed on either side of green part, medially weakly 3-ribbed, ovate to elliptic ovate, (2.4–)2.8–3.3(–3.6) × 1.6–2 mm, apex acute, mucronulate from excurrent midrib.
anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm.
anthers (0.3–)0.4–0.6 mm;
styles 0.4–0.6(–0.8) mm;
stigmas 1–1.4 mm.
black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate.
brown, stipitate, ellipsoid, (1.6–)1.8–2 × (0.7–) 0.8–0.9 mm, stipe 0.1–0.2 × 0.2 mm, apex ± truncate, apiculate, surfaces puncticulate to glabrous.
Cyperus rotundus
Cyperus dentoniae
Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations.
Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus.
Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Cyperus dentoniae was not treated in G. Kükenthal’s monograph (1935–1936), apparently because he did not see any materials of the species. The plant has been treated as “C. asper (Liebmann) O’Neill,” a name based on Mariscus asper Liebmann and a synonym of C. mutisii (A. B. Ayers 1946).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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