Cyperus rotundus |
Cyperus |
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chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
cyperus, flatsedge, galingale, nutsedge, umbrella-sedge |
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Habit | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. | Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous. | ||||||||||||||||
Culms | trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles. |
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Leaves | V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
usually basal; ligules absent; blades keeled abaxially, flat, V-, or inversely W-shaped in cross section. |
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Inflorescences | spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet; spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts; involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike. |
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Spikelets | (2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous). |
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Flowers | anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings; perianth absent; stamens 1–3; styles linear, 2–3-fid, base deciduous or persistent; stigmas 2–3. |
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Achenes | black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
biconvex, flattened, or trigonous. |
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Cyperus rotundus |
Cyperus |
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Phenology | Fruiting summer–fall. | |||||||||||||||||
Habitat | Croplands, disturbed soils usually | |||||||||||||||||
Elevation | 0–400 m (0–1300 ft) | |||||||||||||||||
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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Pantemperate and tropical |
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Discussion | Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 600 (96 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 169. | FNA vol. 23, p. 141. | ||||||||||||||||
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae | ||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754) | ||||||||||||||||
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