Cyperus retrorsus |
Cyperus rotundus |
|
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pine barren flatsedge |
chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
|
Habit | Herbs, perennial, cespitose, shortly rhizomatous. | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. |
Culms | trigonous, 20–50(–85) cm × 1–2(–4) mm, glabrous. |
trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
Leaves | 3–6(–10), 10–40(–55) cm × 1.5–4(–5) mm, V- to inversely W-shaped. |
V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
Inflorescence(s) | spike 1, densely oblong-ovoid often with small basal branches, 8–16 × 4–12(–16) mm; rays 4–8(–11), 0.5–3.5(–8.5) cm; bracts (3–)4–6(–10), ascending at (30–)45°, flat to broadly V-shaped, (4–)14–30(–55) cm × 1–5 mm, scabridulous along margins only; rachilla deciduous, wings persistent, 0.2–0.3 mm wide. |
spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
Spikelets | 40–120, oblong-lanceoloid, subterete, 2.2–4(–4.5) × 0.8–1.6 mm; distal spikelet spreading or ascending; floral scales persistent, 2–5, appressed, brown to stramineous, or fuscous and lustrous, medially greenish and 1(–3)-ribbed, laterally greenish to yellowish white and 3–4-ribbed, oblong to oblong-lanceolate, 1.8–2.5 × 1–1.5 mm, subcoriacous. |
(2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
Flowers | anthers 0.3–0.5 mm; styles 0.5–0.8 mm; stigmas 0.6–0.9 mm. |
anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
Achenes | brown, oblong-ellipsoid, 1.2–1.7 × 0.5–0.6 mm, base cuneate, apex subacute, surfaces papillose. |
black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
Cyperus retrorsus |
Cyperus rotundus |
|
Phenology | Fruiting summer (Jun–Aug). | Fruiting summer–fall. |
Habitat | Open woods and thickets in moist to dry, sandy soils | Croplands, disturbed soils usually |
Elevation | 0–1000 m (0–3300 ft) | 0–400 m (0–1300 ft) |
Distribution |
AL; AR; DC; DE; FL; GA; LA; MD; MO; MS; NC; NJ; NY; OK; SC; TN; TX; VA; Mexico (Tamaulipas)
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AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
|
Discussion | Cyperus retrorsus was reported in Indiana (M. L. Horvat 1941); we have been unable to confirm the report. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 187. | FNA vol. 23, p. 169. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | Mariscus cylindricus, C. cylindricus, Mariscus retrorsus | |
Name authority | Chapman: Bot. Gaz. 3: 17. (1878) | Linnaeus: Sp. Pl. 1: 45. (1753) |
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