Cyperus reflexus |
Cyperus tetragonus |
|
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bentawn flatsedge |
fourangle flatsedge |
|
Habit | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. | Herbs, perennial, cespitose, rhizomatous. |
Culms | roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
trigonous, (20–) 30–80 cm × 1–2.5 mm, glabrous or occasionally with scattered prickles on distal angles. |
Leaves | 1–6, 10–40 cm, glabrous or essentially so. |
9–11, flat to slightly V-shaped, 25–80 cm × 3.5–10 mm. |
Inflorescences | heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
spikes (1–)3–6, loosely cylindric or oblong with parallel sides, 1.5–4 cm × 10–17 mm; rays 6–12, 2–25 cm, glabrous or rarely scabridulous proximal to spikes; bracts 6–11, ascending at 45°, flat, (1–)10–20(–26) cm × 1–10 mm; rachilla deciduous, wings persistent, 0.4–0.5 mm wide. |
Spikelets | 25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
10–60, narrowly ellipsoid, quadrangular, 5–8 × 1.5–2 mm; floral scales persistent, (2–)3–7, appressed, clear at maturity, laterally reddish to purplish brown, medially greenish, often erose at maturity, laterally 3–5-ribbed, medially 3-ribbed, ovate-elliptic, 2.5–3.2 × 1.8–2.1 mm, apex obtuse. |
Flowers | stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
anthers 0.8–1 mm; styles 0.3–0.5 mm; stigmas 2–2.5 mm. |
Achenes | reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
brown to reddish brown, sessile, oblong-ovoid to broadly ellipsoid, slightly dorsiventrally flattened, 1.8–2 × 0.7–0.8 mm, apex obtuse, apiculate, surfaces papillose to puncticulate. |
Cyperus reflexus |
Cyperus tetragonus |
|
Phenology | Fruiting late spring–fall (May–Oct). | Fruiting summer–fall (Jun–Nov). |
Habitat | Muddy soils or shallow waters | Open woods, thickets, barrier islands |
Elevation | 0–200 m (0–700 ft) | 0–1000 m (0–3300 ft) |
Distribution |
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
AL; AZ; FL; GA; MS; NC; NM
|
Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The status of Cyperus pringlei from the southwestern United States requires further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 189. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus | C. pringlei, C. tetragonus var. pringlei, Mariscus tetragonus |
Name authority | Vahl: Enum. Pl. 2: 299. (1805) | Elliott: Sketch Bot. S. Carolina 1: 71. (1821) |
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