Cyperus reflexus |
Cyperus squarrosus |
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bentawn flatsedge |
awn cyperus, awn flatsedge, bearded flatsedge |
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Habit | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. | Herbs, annual, cespitose, with fibrous roots. |
Culms | roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
1–20, trigonous, 2–16 cm × 0.5–0.8 mm, glabrous. |
Leaves | 1–6, 10–40 cm, glabrous or essentially so. |
1–3, flat to V-shaped, (1–)5–10(–15) × 0.5–2.5 mm. |
Inflorescences | heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
spike 1, loosely to densely ovoid to oblong, 6–20(–40) × 9–15(–20) mm; rays absent or 1–3(–6), 4–40 mm; bracts (1–)2–4, longest ± erect, V-shaped, 1–15 cm × 0.5–3 mm; rachilla ± deciduous, wingless. |
Spikelets | 25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
(2–)6–20(–40), greenish to reddish brown, somewhat compressed, ovoid-lanceoloid to oblong, 2.5–10(–20) × 1.3–2.2 mm (excluding awns); floral scales deciduous, (4–)10–20(–34), greenish to stramineous or brownish red laterally, greenish medially, (5–)7–9(–11)-ribbed nearly to margins, oblong-lanceolate, (1.2–)1.3–1.8(–2.2) × (0.5–)0.7–0.8(–1) mm, apex cuspidate, excurved awn additional 0.5–1(–1.3) mm. |
Flowers | stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
stamen 1, filaments 1.5 mm; anthers oblong, 0.2–0.3 mm, connective apex reddish, minute; styles 0.3–0.5 mm; stigmas 0.4–0.7 mm. |
Achenes | reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
light brown to nearly black, stipitate, obovoid (occasionally linear-spatulate or linear oblong, infrequently constricted in middle), 0.7–0.8(–1.1) × (0.2–)0.3–0.4(–0.5) mm, stipe 0.05–0.1 × 0.1 mm, apex truncate, apiculate, surfaces papillose. |
Cyperus reflexus |
Cyperus squarrosus |
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Phenology | Fruiting late spring–fall (May–Oct). | Fruiting summer. |
Habitat | Muddy soils or shallow waters | Moist, disturbed soils, gravelly roadsides, flood plains, edges of puddles, muddy places |
Elevation | 0–200 m (0–700 ft) | 0–2500 m (0–8200 ft) |
Distribution |
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; ON; QC; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
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Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus squarrosus can be recognized by its small size and annual habit combined with its oblong-lanceolate floral scales bearing five to eleven conspicuous ribs and excurved awns. Some collections have been misidentified as C. acuminatus, an annual species of subg. Pycnostachys that has ovate-lanceolate, three-ribbed floral scales and digitately clustered spikelets. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 165. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus | Chlorocyperus inflexus, C. aristatus, C. aristatus var. inflexus, C. aristatus var. runyonii, C. inflexus, Dichostylis aristata, Mariscus squarrosus |
Name authority | Vahl: Enum. Pl. 2: 299. (1805) | Linnaeus: Cent. Pl. II, 6. (1756) |
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