Cyperus reflexus |
Cyperus schweinitzii |
|
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bentawn flatsedge |
sand flatsedge, Schweinitz's flatsedge |
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Habit | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. | Herbs, perennial, rhizomatous. |
Culms | roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
trigonous, (10–)20–50 cm × (0.6–)1–2 mm, glabrous or more often scabridulous, either in distal 1/2 or over entire length; basal sheaths, 1–3, light brown, 1–4 cm, papery. |
Leaves | 1–6, 10–40 cm, glabrous or essentially so. |
flat to V-shaped, (2–)20–35 cm × 2–6 mm, margins and keel minutely scabridulous; proximal leaves bladeless. |
Inflorescences | heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
spikes ± umbellate to slenderly (infrequently broadly) ovoid, 10–25 × 7–16 mm; rays 3–5, 1.5–8(–15) cm; bracts 3–7, erect (or declined at most 25–30° from vertical), (2.5–)8–20(–25) cm × (0.5–)1.5–6 mm; rachilla ± deciduous, wingless. |
Spikelets | 25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
(1–)5–10(–20), oblong to linear-oblong, quadrangular, 7–10(–18) × (2.8–)3.2–4(–4.5) mm; floral scales deciduous, (1–)5–10(–14), spreading, laterally stramineous or dull whitish, sometimes also red-spotted, medially green, laterally 2–3-ribbed, medially 3-ribbed, broadly ovate to oblong-ovate, (2.3–)2.8–3.2 × (2–)2.4–3.2 mm, apex broadly rounded, often clear-erose, cusp 0.1–0.4 mm in proximal scales, 0.3–1 mm in distal scales. |
Flowers | stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
anthers 0.8–1.4 mm; styles 0.1–0.3 mm; stigmas 1.5–3 mm. |
Achenes | reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
light to dark brown or nearly black, broadly ellipsoid, infrequently obovoid, 2–2.4 × 0.9–1.4 mm, base cuneate to ± stipelike, apex obtuse to broadly rounded, surfaces papillose. |
2n | = 166. |
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Cyperus reflexus |
Cyperus schweinitzii |
|
Phenology | Fruiting late spring–fall (May–Oct). | Fruiting late spring–early summer. |
Habitat | Muddy soils or shallow waters | Riverbanks, sand bars, lakeshores, sand dunes, sandy openings in woods |
Elevation | 0–200 m (0–700 ft) | 0–1000 m (0–3300 ft) |
Distribution |
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
CO; IA; IL; KY; MA; MI; MN; MO; MT; NJ; NM; NY; OH; OK; PA; SD; TX; UT; WA; WI; WY; AB; MB; ON; QC; SK; Mexico (Coahuila)
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Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus schweinitzii is introduced, but not naturalized, in Massachusetts and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 174. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus | C. alterniflorus |
Name authority | Vahl: Enum. Pl. 2: 299. (1805) | Torrey: Ann. Lyceum Nat. Hist. New York 3: 276. (1836) |
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