FNA: Cyperus reflexus vs. Cyperus sanguinolentus

	Cyperus reflexus	Cyperus sanguinolentus
	bentawn flatsedge	purpleglume flatsedge
Habit	Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide.	Herbs, perennial; rhizomes ± horizontal, to 12 cm × 0.8–1.2 mm.
Culms	roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.	± terete to roundly trigonous, 3–25(–60) cm × 0.3–2 mm, glabrous.
Leaves	1–6, 10–40 cm, glabrous or essentially so.	1–3, V-shaped, 1–8(–15) cm × 1–2 mm.
Inflorescences	heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.	spike 1, loosely ovoid, 7–14 × 8–20 mm; rays (0–)1–4, to 2(–3) cm; bracts 2–4, horizontal to ascending at 30(–45)°, V-shaped to flat, 1–8(–18) cm × 0.5–1.5(–2) mm.
Spikelets	25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).	3–5(–14), oblong to linear-lanceoloid, 8–18 × 2–2.6(–3) mm; floral scales (6–)10–26(–32), laterally clear to light brown, sometimes with purplish margins, medially light brown, 2-keeled, laterally ribless, medially 2–3-ribbed, oblong to ovate, 1.9–2.2(–2.7) × 1.8–2.3 mm, apex obtuse.
Flowers	stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm.	stamens 3; anthers ellipsoid, 0.3–0.6 mm, connective not prolonged; styles 0.6–1 mm; stigmas 1–1.5 mm.
Achenes	reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.	brown, ± stipitate, obovoid to ovoid, 1–1.4 × 0.6–0.8 mm, apex truncate, apiculate, surfaces minutely punctate.

	Cyperus reflexus	Cyperus sanguinolentus
Phenology	Fruiting late spring–fall (May–Oct).	Fruiting late summer–fall.
Habitat	Muddy soils or shallow waters	Emergent shorelines, ditches
Elevation	0–200 m (0–700 ft)	20 m (100 ft)
Distribution	FL; LA; OK; TX; Mexico; Central America (Costa Rica) [BONAP county map]	AL; GA; LA; MS; Asia; Africa [Introduced in North America] [BONAP county map]
Discussion	Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Cyperus sanguinolentus is the only rhizomatous, distigmatic Cyperus in the eastern United States. An early collection from Louisiana was described as C. louisianensis, supposedly endemic. Recent field work and morphometric studies convincingly showed it to be an introduction of the Asian C. sanguinolentus) (J. R. Carter and C. T. Bryson 2001. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23.	FNA vol. 23, p. 161.
Parent taxa	Cyperaceae > Cyperus > subg. Pycnostachys	Cyperaceae > Cyperus > subg. Pycreus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus	C. louisianensis, Pycreus sanguinolentus
Name authority	Vahl: Enum. Pl. 2: 299. (1805)	Vahl: Enum. Pl. 2: 351. (1805)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus sanguinolentus

bentawn flatsedge

purpleglume flatsedge

Habit

Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide.

Herbs, perennial; rhizomes ± horizontal, to 12 cm × 0.8–1.2 mm.

Culms

roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.

± terete to roundly trigonous, 3–25(–60) cm × 0.3–2 mm, glabrous.

Leaves

1–6, 10–40 cm, glabrous or essentially so.

1–3, V-shaped, 1–8(–15) cm × 1–2 mm.

Inflorescences

heads ovoid to pyramidal, 9–15 mm diam.;

rays 0 or 3–6, 0.5–4 cm, glabrous;

bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.

spike 1, loosely ovoid, 7–14 × 8–20 mm;

rays (0–)1–4, to 2(–3) cm;

bracts 2–4, horizontal to ascending at 30(–45)°, V-shaped to flat, 1–8(–18) cm × 0.5–1.5(–2) mm.

Spikelets

25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm;

floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).

3–5(–14), oblong to linear-lanceoloid, 8–18 × 2–2.6(–3) mm;

floral scales (6–)10–26(–32), laterally clear to light brown, sometimes with purplish margins, medially light brown, 2-keeled, laterally ribless, medially 2–3-ribbed, oblong to ovate, 1.9–2.2(–2.7) × 1.8–2.3 mm, apex obtuse.

Flowers

stamen 1;

anthers 0.6–1 mm;

styles 0.3–0.5 mm;

stigmas 0.3–0.6 mm.

stamens 3;

anthers ellipsoid, 0.3–0.6 mm, connective not prolonged;

styles 0.6–1 mm;

stigmas 1–1.5 mm.

Achenes

reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.

brown, ± stipitate, obovoid to ovoid, 1–1.4 × 0.6–0.8 mm, apex truncate, apiculate, surfaces minutely punctate.

Cyperus reflexus

Cyperus sanguinolentus

Phenology

Fruiting late spring–fall (May–Oct).

Fruiting late summer–fall.

Habitat

Muddy soils or shallow waters

Emergent shorelines, ditches

Elevation

0–200 m (0–700 ft)

20 m (100 ft)

Distribution

FL; LA; OK; TX; Mexico; Central America (Costa Rica)

[BONAP county map]

AL; GA; LA; MS; Asia; Africa [Introduced in North America]

[BONAP county map]

Discussion

Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus sanguinolentus is the only rhizomatous, distigmatic Cyperus in the eastern United States. An early collection from Louisiana was described as C. louisianensis, supposedly endemic. Recent field work and morphometric studies convincingly showed it to be an introduction of the Asian C. sanguinolentus) (J. R. Carter and C. T. Bryson 2001.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23.

FNA vol. 23, p. 161.

Parent taxa

Cyperaceae > Cyperus > subg. Pycnostachys

Cyperaceae > Cyperus > subg. Pycreus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus

C. louisianensis, Pycreus sanguinolentus

Name authority

Vahl: Enum. Pl. 2: 299. (1805)

Vahl: Enum. Pl. 2: 351. (1805)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Cyperus reflexus

Cyperus sanguinolentus

Cyperus reflexus

Cyperus sanguinolentus