Cyperus reflexus |
Cyperus refractus |
|
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bentawn flatsedge |
reflexed flatsedge |
|
Habit | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. | Herbs, perennial, cespitose, with inconspicuous, short rhizomes scarcely visible between culm bases. |
Culms | roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
basally cormlike, trigonous, 30–100 cm × 2–4 mm, glabrous. |
Leaves | 1–6, 10–40 cm, glabrous or essentially so. |
V-shaped to flat, 20–70 cm × 3–8 mm, adaxial surface, margins minutely scabridulous. |
Inflorescences | heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
spikes loose, broadly ellipsoid to ovoid, 4–7 × 2–6 cm; rays 6–13(–16), 4–25 cm, glabrous; rachis 1.5–3 cm; bracts 3–6, horizontal to ascending at 30°, flat, 5–35 cm × 2.5–6 mm; rachilla persistent, wings 0.4–0.5 mm wide. |
Spikelets | 25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
(13–)20–60(–74), linear, ± terete, (11–)13–25(–28) × 1.2–1.5 mm; distal spikelets spreading or ascending; floral scales persistent, (2–)3–6, stramineous, red-purple striate, laterally 3–5-ribbed, ovate-lanceolate, 4.5–5.3 × 1.2–1.5 mm. |
Flowers | stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
anthers 1–1.5 mm; styles 1.5 mm; stigmas 1.5 mm. |
Achenes | reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
brown to purplish brown, oblong-linear (5–6 times long as wide), 2.6–3 × 0.5–0.6 mm, base cuneate, apex obtuse, surfaces puncticulate. |
Cyperus reflexus |
Cyperus refractus |
|
Phenology | Fruiting late spring–fall (May–Oct). | Fruiting summer (Jul–Aug). |
Habitat | Muddy soils or shallow waters | Sandy riverbanks, gravelly roadsides, pastures, disturbed places |
Elevation | 0–200 m (0–700 ft) | 20–300 m (100–1000 ft) |
Distribution |
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
AL; AR; DC; GA; MD; MO; NC; PA; SC; TN; VA; WV |
Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus refractus is often mistaken for C. strigosus, which has shorter anthers (0.4–0.5 mm) and spreading, ± deciduous floral scales. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 182. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus | |
Name authority | Vahl: Enum. Pl. 2: 299. (1805) | Engelmann ex Boeckeler: Linnaea 36: 369. 1870, based on Cyperus refrofractus Engelmann ex Steudel, Syn. Pl. Glumac. 2: 47. 1855, not (Linnaeus) Torrey (1848) |
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