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bentawn flatsedge

pond flatsedge

Habit Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. Herbs, perennial, cespitose; rhizomes oblique.
Culms

roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.

roundly trigonous, 20–50(–80) cm × 1–3 mm, glabrous.

Leaves

1–6, 10–40 cm, glabrous or essentially so.

3–11, V-shaped, 12–40 cm.

Inflorescences

heads ovoid to pyramidal, 9–15 mm diam.;

rays 0 or 3–6, 0.5–4 cm, glabrous;

bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.

heads loosely to rather densely hemispheric, 15–30 mm diam.;

rays 5–10, 2–12 cm; 2d order rays occasionally produced, 5–25 mm;

bracts 4–8, approximately horizontal to ascending at 30°, V-shaped, 10–36 cm × 1.5–4 mm, margins glabrous.

Spikelets

25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm;

floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).

(4–)10–30, greenish, grayish, or stramineous, oblong to linear-oblong, compressed, (5–)7–10(–35) × 2–3 mm;

floral scales 10–25(–85), laterally yellowish brown, medially yellowish or brownish, laterally ribless, medially 2-ribbed with groove between ribs, broadly ovate (widest in proximal 1/2), (1.2–)1.5–2 × 1.2–2 mm, apex obtuse (declined (45–)60–90º from rachilla).

Flowers

stamen 1;

anthers 0.6–1 mm;

styles 0.3–0.5 mm;

stigmas 0.3–0.6 mm.

stamen 1;

anthers oblong, 0.8–1.2 mm, connective not prolonged;

styles basally thickened, confluent with summit of achene, 0.6–1.2 mm;

stigmas 0.3–0.6 mm.

Achenes

reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.

reddish brown to dark purplish brown, dull or more often iridescent, stipitate, broadly ellipsoid, 1–1.5 mm × 0.4–0.6(–0.7) mm, base cuneate to ± attenuate, stipe 0.1–0.2 mm, apex acute, beak 0.1–0.5 mm, surfaces reticulate with network of low ridges.

Cyperus reflexus

Cyperus ochraceus

Phenology Fruiting late spring–fall (May–Oct). Fruiting summer.
Habitat Muddy soils or shallow waters Damp, disturbed soils, ditches, marshes
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
FL; LA; OK; TX; Mexico; Central America (Costa Rica)
[BONAP county map]
from FNA
FL; GA; LA; MS; PA; TX; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Pennsylvania, Cyperus ochraceus has been recorded in areas where ballast has been discarded.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 155.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Pycnostachys
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus
Name authority Vahl: Enum. Pl. 2: 299. (1805) Vahl: Enum. Pl. 2: 325. (1805)
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