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bentawn flatsedge

black flatsedge, brown cyperus

Habit Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. Herbs, perennial, rhizomatous.
Culms

roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.

solitary or close together, trigonous, 5–40(–95) cm, glabrous.

Leaves

1–6, 10–40 cm, glabrous or essentially so.

(2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm.

Inflorescences

heads ovoid to pyramidal, 9–15 mm diam.;

rays 0 or 3–6, 0.5–4 cm, glabrous;

bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.

spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.;

rachis 1–2 mm, or absent;

rays 1–2, 3–45 mm;

if rays absent, inflorescence sessile, dense, 6–20 mm diam;

bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm.

Spikelets

25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm;

floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).

(3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm;

floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse.

Flowers

stamen 1;

anthers 0.6–1 mm;

styles 0.3–0.5 mm;

stigmas 0.3–0.6 mm.

stamens 2;

anthers 0.6–0.8 mm, connectives not prolonged;

styles 0.4–0.6 mm;

stigmas 0.7–2.1 mm.

Achenes

reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.

brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate.

Cyperus reflexus

Cyperus niger

Phenology Fruiting late spring–fall (May–Oct). Fruiting summer.
Habitat Muddy soils or shallow waters Swamps, ditches, wet pastures, seeps in montane forests
Elevation 0–200 m (0–700 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
FL; LA; OK; TX; Mexico; Central America (Costa Rica)
[BONAP county map]
from FNA
AZ; CA; NM; OK; TX; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Pycreus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus
Name authority Vahl: Enum. Pl. 2: 299. (1805) Ruiz & Pavón: Fl. Peruv. 1: 47. (1798)
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