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bentawn flatsedge

bristly flatsedge

Habit Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. Herbs, perennial, cespitose, with well-developed rhizomes.
Culms

roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.

trigonous, 30–100 cm × 2–4 mm, basally cormlike, glabrous.

Leaves

1–6, 10–40 cm, glabrous or essentially so.

flat to broadly V-shaped, 20–70 cm × 4–6 mm, glabrous except for marginal prickles.

Inflorescences

heads ovoid to pyramidal, 9–15 mm diam.;

rays 0 or 3–6, 0.5–4 cm, glabrous;

bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.

spikes loosely to densely ovoid, oblong (2 times long as wide), 10–12 × 6–9 mm;

rays 6–11, 1–16 cm, glabrous;

bracts 5–10, ascending at 30–45°, flat, 6–25 cm × 3–6 mm;

rachilla persistent, wings 0.5 mm wide, covering nearly entire length of achene.

Spikelets

25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm;

floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).

(20–)40–100(–120), proximal spikelets reflexed somewhat, distal ones divaricate, ± terete, lanceoloid, 6–6.8 × 1–1.4 mm;

floral scales persistent, 1–2(–3), appressed, golden brown, lanceolate, laterally 5–6-ribbed, 3.8–4.9 × 1.4–1.6 mm, apex mucronate, mucro at most 0.3 mm.

Flowers

stamen 1;

anthers 0.6–1 mm;

styles 0.3–0.5 mm;

stigmas 0.3–0.6 mm.

anthers 1–1.3 mm;

styles 0.8–1 mm;

stigmas 2–3 mm.

Achenes

reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.

brown, sessile, linear, 2.5–2.8 × 0.5–0.7 mm, apex obtuse, apiculate, surfaces muriculate.

Cyperus reflexus

Cyperus hystricinus

Phenology Fruiting late spring–fall (May–Oct). Fruiting late summer–early fall (Jul–Sep).
Habitat Muddy soils or shallow waters Xeric, sandy soils of sand hills and pine barrens
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
FL; LA; OK; TX; Mexico; Central America (Costa Rica)
[BONAP county map]
from FNA
AL; AR; DC; DE; GA; LA; MD; MO; MS; NC; NJ; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus hystricinus resembles C. plukenetii and C. retrofractus; it can be readily distinguished from both by its glabrous culms. Overly mature specimens of C. lancastriensis are frequently misidentified as C. hystricinus; C. hystricinus may be confirmed by its narrow, nearly glabrous leaves and bracts, golden brown spikelets, longer, narrower achenes, and elongated rhizome internodes (to 15 mm vs. 5 mm in 76. C. lancastriensis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus C. retrofractus var. hystricinus
Name authority Vahl: Enum. Pl. 2: 299. (1805) Fernald: Rhodora 8: 127. (1906)
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