Cyperus reflexus |
Cyperus erythrorhizos |
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bentawn flatsedge |
red-root cyperus, red-root flat sedge, redroot nutgrass |
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Habit | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. | Herbs, annual, cespitose. |
Culms | roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
trigonous to roundly trigonous, (0.5–)5–25(–105) cm × 1–2.5(–7.5) mm, glabrous. |
Leaves | 1–6, 10–40 cm, glabrous or essentially so. |
flat to inversely W-shaped, 5–25(–90) cm × 2–5(–11) mm. |
Inflorescences | heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
spikes 1–3(–6), rather densely cylindric-ovoid, (4–)10–30(–45) × (6–)10–16(–23) mm; rays 2–6(–12), 1–8(–28) cm; bracts (3–)5–7(–11), horizontal to ascending at 30°, inversely W-shaped, (3–)10–20(–70) cm × 1–3(–12) mm; rachilla persistent, at maturity becoming laterally free, remaining firmly attached proximally, wings 0.3(–0.4) mm wide. |
Spikelets | 25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
(20–)40–80, linear, quadrangular, 3–8(–11) × 1–1.5 mm; floral scales deciduous, 6–16(–30), appressed, laterally light brown with reddish speckles, medially greenish, laterally ribless, medially 3-ribbed, oblong-ovate to obovate, quadrangular to subterete, 1.3–1.5 × 0.8–1.2 mm, apex obtuse, mucronulate. |
Flowers | stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
anthers 0.2–0.3 mm, connectives 0.1 mm; styles 0.7–0.8 mm; stigmas 0.4 mm. |
Achenes | reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
light grayish to brown, sessile, ovoid, (0.4–)0.7–1 × 0.4–0.6 mm, apex rounded, apiculate, surfaces glabrous. |
Cyperus reflexus |
Cyperus erythrorhizos |
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Phenology | Fruiting late spring–fall (May–Oct). | Fruiting summer. |
Habitat | Muddy soils or shallow waters | Emergent shorelines |
Elevation | 0–200 m (0–700 ft) | 0–1500 m (0–4900 ft) |
Distribution |
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
AL; AR; AZ; CA; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; WA; WI; WV; MB; ON; Mexico (Baja California)
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Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 23. | FNA vol. 23, p. 172. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus | C. cupreus, C. erythrorhizos var. cupreus, C. occidentalis, C. washingtonensis |
Name authority | Vahl: Enum. Pl. 2: 299. (1805) | Muhlenberg: Descr. Gram., 20. (1817) |
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