FNA: Cyperus reflexus vs. Cyperus drummondii

	Cyperus reflexus	Cyperus drummondii
	bentawn flatsedge	Drummond's sedge
Habit	Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide.	Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent.
Culms	roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.	trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles.
Leaves	1–6, 10–40 cm, glabrous or essentially so.	4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm.
Inflorescences	heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.	heads digitate, globose, 1–1.7 cm diam.; rays 3–5, 1–7 cm; 2d order rays absent; bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent.
Spikelets	25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).	10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm; floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous.
Flowers	stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm.	stamens 1–2; anthers 0.8–1.2 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm.
Achenes	reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.	brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous.

	Cyperus reflexus	Cyperus drummondii
Phenology	Fruiting late spring–fall (May–Oct).	Fruiting late spring–summer.
Habitat	Muddy soils or shallow waters	Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies
Elevation	0–200 m (0–700 ft)	0–200 m (0–700 ft)
Distribution	FL; LA; OK; TX; Mexico; Central America (Costa Rica) [BONAP county map]	AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies [BONAP county map]
Discussion	Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate. Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23.	FNA vol. 23.
Parent taxa	Cyperaceae > Cyperus > subg. Pycnostachys	Cyperaceae > Cyperus > subg. Pycnostachys
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus	C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus
Name authority	Vahl: Enum. Pl. 2: 299. (1805)	Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus drummondii

bentawn flatsedge

Drummond's sedge

Habit

Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide.

Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent.

Culms

roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.

trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles.

Leaves

1–6, 10–40 cm, glabrous or essentially so.

4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm.

Inflorescences

heads ovoid to pyramidal, 9–15 mm diam.;

rays 0 or 3–6, 0.5–4 cm, glabrous;

bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.

heads digitate, globose, 1–1.7 cm diam.;

rays 3–5, 1–7 cm; 2d order rays absent;

bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent.

Spikelets

25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm;

floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).

10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm;

floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous.

Flowers

stamen 1;

anthers 0.6–1 mm;

styles 0.3–0.5 mm;

stigmas 0.3–0.6 mm.

stamens 1–2;

anthers 0.8–1.2 mm;

styles 0.8–1 mm;

stigmas 0.6–0.8 mm.

Achenes

reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.

brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous.

Cyperus reflexus

Cyperus drummondii

Phenology

Fruiting late spring–fall (May–Oct).

Fruiting late spring–summer.

Habitat

Muddy soils or shallow waters

Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies

Elevation

0–200 m (0–700 ft)

Distribution

FL; LA; OK; TX; Mexico; Central America (Costa Rica)

[BONAP county map]

AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies

[BONAP county map]

Discussion

Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate.

Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23.

Parent taxa

Cyperaceae > Cyperus > subg. Pycnostachys

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus

C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus

Name authority

Vahl: Enum. Pl. 2: 299. (1805)

Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Cyperus reflexus

Cyperus drummondii

Cyperus reflexus

Cyperus drummondii