Cyperus reflexus |
Cyperus |
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bentawn flatsedge |
cyperus, flatsedge, galingale, nutsedge, umbrella-sedge |
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Habit | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. | Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous. | ||||||||||||||||
Culms | roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles. |
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Leaves | 1–6, 10–40 cm, glabrous or essentially so. |
usually basal; ligules absent; blades keeled abaxially, flat, V-, or inversely W-shaped in cross section. |
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Inflorescences | heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet; spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts; involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike. |
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Spikelets | 25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous). |
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Flowers | stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings; perianth absent; stamens 1–3; styles linear, 2–3-fid, base deciduous or persistent; stigmas 2–3. |
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Achenes | reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
biconvex, flattened, or trigonous. |
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Cyperus reflexus |
Cyperus |
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Phenology | Fruiting late spring–fall (May–Oct). | |||||||||||||||||
Habitat | Muddy soils or shallow waters | |||||||||||||||||
Elevation | 0–200 m (0–700 ft) | |||||||||||||||||
Distribution |
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
Pantemperate and tropical |
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Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 600 (96 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23. | FNA vol. 23, p. 141. | ||||||||||||||||
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae | ||||||||||||||||
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Subordinate taxa | ||||||||||||||||||
Synonyms | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus | |||||||||||||||||
Name authority | Vahl: Enum. Pl. 2: 299. (1805) | Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754) | ||||||||||||||||
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