FNA: Cyperus reflexus vs. Cyperus hypopitys

	Cyperus reflexus	Cyperus hypopitys
	bentawn flatsedge	pine flatsedge, pinewoods sedge
Habit	Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide.	Herbs, perennial, rhizomatous, tending to form small clumps by basal offshoots.
Culms	roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.	triquetrous, bulbous, thickened, (20–)30–80(–90) cm × 0.7–1.2 mm, glabrous proximally, becoming trigonous strongly scabridulous on distal angles, clothed with persistent reddish fibers at base.
Leaves	1–6, 10–40 cm, glabrous or essentially so.	flat to V-shaped, 20–45 cm × 2–4 mm.
Inflorescences	heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.	spike 1, loosely ovoid to ovoid-ellipsoid, (1.5–)2.5–7 × (1–)1.5–3.5 cm; rays (1–)2–4, to 6 cm; 2d order rays rarely present, to 2 cm; 3d order rays absent; bracts (2–) 3, ± horizontal, 1–15(–21) cm × 0.4–3.5 mm; rachilla deciduous, wings hyaline, 0.3–0.4 mm wide.
Spikelets	25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).	(3–)8–26, reddish to reddish brown, linear, compressed-quadrangular, 8–18 × (2.4–)2.8–4 mm; floral scales deciduous, 6–16, laterally reddish, medially green, occasionally red-glandular, laterally strongly 3(–4)-ribbed, medially punctate, 1–3-ribbed, ovate-lanceolate, (2.6–)2.8–3.3 × (1.4–)1.6–1.8(–2) mm, apex weakly 3-dentate, mucronulate, with tuft of crystalline prickles, medially glabrous.
Flowers	stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm.	anthers 0.8–1.1 mm; styles 1–1.4 mm; stigmas 1–1.4 mm.
Achenes	reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.	brownish, stipitate, ellipsoid, 1.6–2 × (0.7–)0.9–1 mm, base whitish, stipe to 0.1–0.2 mm, apex obtuse, slightly apiculate, surfaces papillose.

	Cyperus reflexus	Cyperus hypopitys
Phenology	Fruiting late spring–fall (May–Oct).	Fruiting late summer–early fall.
Habitat	Muddy soils or shallow waters	Canyons in pine forests
Elevation	0–200 m (0–700 ft)	500–1500 m (1600–4900 ft)
Distribution	FL; LA; OK; TX; Mexico; Central America (Costa Rica) [BONAP county map]	AZ; Mexico (Chihuahua, Durango, Sinaloa, Sonora) [BONAP county map]
Discussion	Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Of conservation concern. Cyperus hypopitys can be recognized by the combination of reddish floral scales and sharp-angled culms. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23.	FNA vol. 23, p. 178.
Parent taxa	Cyperaceae > Cyperus > subg. Pycnostachys	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus
Name authority	Vahl: Enum. Pl. 2: 299. (1805)	G. C. Tucker: Syst. Bot. Monogr. 43: 129, fig. 54. (1994)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus hypopitys

bentawn flatsedge

pine flatsedge, pinewoods sedge

Habit

Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide.

Herbs, perennial, rhizomatous, tending to form small clumps by basal offshoots.

Culms

roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous.

triquetrous, bulbous, thickened, (20–)30–80(–90) cm × 0.7–1.2 mm, glabrous proximally, becoming trigonous strongly scabridulous on distal angles, clothed with persistent reddish fibers at base.

Leaves

1–6, 10–40 cm, glabrous or essentially so.

flat to V-shaped, 20–45 cm × 2–4 mm.

Inflorescences

heads ovoid to pyramidal, 9–15 mm diam.;

rays 0 or 3–6, 0.5–4 cm, glabrous;

bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm.

spike 1, loosely ovoid to ovoid-ellipsoid, (1.5–)2.5–7 × (1–)1.5–3.5 cm;

rays (1–)2–4, to 6 cm; 2d order rays rarely present, to 2 cm; 3d order rays absent;

bracts (2–) 3, ± horizontal, 1–15(–21) cm × 0.4–3.5 mm;

rachilla deciduous, wings hyaline, 0.3–0.4 mm wide.

Spikelets

25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm;

floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous).

(3–)8–26, reddish to reddish brown, linear, compressed-quadrangular, 8–18 × (2.4–)2.8–4 mm;

floral scales deciduous, 6–16, laterally reddish, medially green, occasionally red-glandular, laterally strongly 3(–4)-ribbed, medially punctate, 1–3-ribbed, ovate-lanceolate, (2.6–)2.8–3.3 × (1.4–)1.6–1.8(–2) mm, apex weakly 3-dentate, mucronulate, with tuft of crystalline prickles, medially glabrous.

Flowers

stamen 1;

anthers 0.6–1 mm;

styles 0.3–0.5 mm;

stigmas 0.3–0.6 mm.

anthers 0.8–1.1 mm;

styles 1–1.4 mm;

stigmas 1–1.4 mm.

Achenes

reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose.

brownish, stipitate, ellipsoid, 1.6–2 × (0.7–)0.9–1 mm, base whitish, stipe to 0.1–0.2 mm, apex obtuse, slightly apiculate, surfaces papillose.

Cyperus reflexus

Cyperus hypopitys

Phenology

Fruiting late spring–fall (May–Oct).

Fruiting late summer–early fall.

Habitat

Muddy soils or shallow waters

Canyons in pine forests

Elevation

0–200 m (0–700 ft)

500–1500 m (1600–4900 ft)

Distribution

FL; LA; OK; TX; Mexico; Central America (Costa Rica)

[BONAP county map]

AZ; Mexico (Chihuahua, Durango, Sinaloa, Sonora)

[BONAP county map]

Discussion

Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Cyperus hypopitys can be recognized by the combination of reddish floral scales and sharp-angled culms.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23.

FNA vol. 23, p. 178.

Parent taxa

Cyperaceae > Cyperus > subg. Pycnostachys

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus

Name authority

Vahl: Enum. Pl. 2: 299. (1805)

G. C. Tucker: Syst. Bot. Monogr. 43: 129, fig. 54. (1994)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Cyperus reflexus

Cyperus hypopitys

Cyperus reflexus

Cyperus hypopitys