FNA: Cyperus prolixus vs. Cyperus fendlerianus

	Cyperus prolixus	Cyperus fendlerianus
	mosquito flatsedge	Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge
Habit	Herbs, perennial; rhizomes 3–20 cm × 1–2 cm, indurate.	Herbs, perennial, rhizomatous.
Culms	stoutly trigonous, thickened, 50–300 cm × 5–30 mm, basally indurate, glabrous.	trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm.
Leaves	V-shaped, with cross ribs, 40–130 cm × 10–15 mm, margins and keel scabrid.	2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm.
Inflorescences	spikes appressed-ascending, narrowly cylindric, (2–)3–45 mm × (4–)7–10 mm; rays (5–)9–12, 2–22(–30) cm; 2d order rays distinctly flattened, 1–10 cm; 3d order rays flattened, 0.5–3 cm; rachis 2.5–4 cm; bracts (6–)9–10, ascending at 45–75°, V- or inversely W-shaped, (5–)15–90 cm × 0.5–20 mm; rachilla persistent, wings 0.2–0.4 mm wide.	spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large; rays absent; bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm; rachilla ± deciduous, wingless.
Spikelets	(4–)10–35, appressed-ascending, compressed, linear, 7–12(–15) × 0.6–1.2 mm; floral scales deciduous, 6–20, marginally clear, laterally brown, medially green, laterally 2–3-ribbed, medially 1-ribbed, oblong-ovate, (3.2–) 3.8–4.3 × 1.2–1.7 mm, apex spreading, acute to obtuse, mucronulate.	(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm; floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous.
Flowers	anthers 0.7–0.9 mm, connective apex bright red, oblong, to 0.1 mm; styles 0.8–1.3 mm; stigmas 2–3.3 mm.	anthers 0.4–0.8 mm; styles 0.7–1.3 mm; stigmas 0.7–1.2 mm.
Achenes	brown, sessile, narrowly ellipsoid, 1.8–2.5 × 0.8–1.3 mm, apex ± acute, not apiculate, surfaces puncticulate.	finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous.

	Cyperus prolixus	Cyperus fendlerianus
Phenology	Fruiting summer.	Fruiting summer.
Habitat	Coastal marshes	Clearings in montane forests
Elevation	0–10 m [0–30 ft]	500–1800 m [1600–5900 ft]
Distribution	LA; Mexico; Central America; South America [BONAP county map]	AZ; CO; NM; TX; WY; Mexico [WildflowerSearch map] [BONAP county map]
Discussion	Cyperus prolixus produces large plants, similar in habit to C. giganteus and C. papyrus but easily distinguished by flattened rays. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23, p. 167.	FNA vol. 23, p. 177.
Parent taxa	Cyperaceae > Cyperus > subg. Cyperus	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	C. amplissimus	Chlorocyperus fendlerianus
Name authority	Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 1: 206. (1816)	Boeckeler: Linnaea 35: 520. (1868)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus fendlerianus

mosquito flatsedge

Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge

Habit

Herbs, perennial; rhizomes 3–20 cm × 1–2 cm, indurate.

Herbs, perennial, rhizomatous.

Culms

stoutly trigonous, thickened, 50–300 cm × 5–30 mm, basally indurate, glabrous.

trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm.

Leaves

V-shaped, with cross ribs, 40–130 cm × 10–15 mm, margins and keel scabrid.

2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm.

Inflorescences

spikes appressed-ascending, narrowly cylindric, (2–)3–45 mm × (4–)7–10 mm;

rays (5–)9–12, 2–22(–30) cm; 2d order rays distinctly flattened, 1–10 cm; 3d order rays flattened, 0.5–3 cm;

rachis 2.5–4 cm;

bracts (6–)9–10, ascending at 45–75°, V- or inversely W-shaped, (5–)15–90 cm × 0.5–20 mm;

rachilla persistent, wings 0.2–0.4 mm wide.

spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large;

rays absent;

bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm;

rachilla ± deciduous, wingless.

Spikelets

(4–)10–35, appressed-ascending, compressed, linear, 7–12(–15) × 0.6–1.2 mm;

floral scales deciduous, 6–20, marginally clear, laterally brown, medially green, laterally 2–3-ribbed, medially 1-ribbed, oblong-ovate, (3.2–) 3.8–4.3 × 1.2–1.7 mm, apex spreading, acute to obtuse, mucronulate.

(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm;

floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous.

Flowers

anthers 0.7–0.9 mm, connective apex bright red, oblong, to 0.1 mm;

styles 0.8–1.3 mm;

stigmas 2–3.3 mm.

anthers 0.4–0.8 mm;

styles 0.7–1.3 mm;

stigmas 0.7–1.2 mm.

Achenes

brown, sessile, narrowly ellipsoid, 1.8–2.5 × 0.8–1.3 mm, apex ± acute, not apiculate, surfaces puncticulate.

finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous.

Cyperus prolixus

Cyperus fendlerianus

Phenology

Fruiting summer.

Habitat

Coastal marshes

Clearings in montane forests

Elevation

0–10 m [0–30 ft]

500–1800 m [1600–5900 ft]

Distribution

LA; Mexico; Central America; South America

[BONAP county map]

AZ; CO; NM; TX; WY; Mexico

[WildflowerSearch map]

[BONAP county map]

Discussion

Cyperus prolixus produces large plants, similar in habit to C. giganteus and C. papyrus but easily distinguished by flattened rays.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23, p. 167.

FNA vol. 23, p. 177.

Parent taxa

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

C. amplissimus

Chlorocyperus fendlerianus

Name authority

Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 1: 206. (1816)

Boeckeler: Linnaea 35: 520. (1868)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Flora of North America species comparison

Cyperus prolixus

Cyperus fendlerianus

Cyperus prolixus

Cyperus fendlerianus