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many-spike flatsedge

cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, perennial (annual?), cespitose. Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

trigonous or ± terete, (3–)20–45(–70) cm × (0.8–) 1.2–2 mm, glabrous.

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

Leaves

(2–) 6–10(–16), flat to V-shaped, (2–) 10–30(–60) cm × 1–4 mm.

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

spike 1, loosely ovoid to densely turbinate, 12–40 × 10–25 mm;

rachis 3–10 mm;

rays 1–8(–12), 0.5–3(–6) cm; 2d order rays absent;

if rays absent, inflorescence single dense cluster of spikelets, 12–40 mm diam.;

bracts (3–)4–6(–7), horizontal to ascending at 45°, flanged V-shaped, (2–)5–15(–23) cm × 1–4 mm.

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

Spikelets

(5–)15–30(–40), spreading to appressed-ascending, linear-lanceoloid to linear, compressed, 8–18(–40) × (1–)1.3–1.6(–2) mm;

floral scales (8–)12–25(–66), laterally stramineous to light brown, medially green, laterally ribless, medially 3–5-ribbed, oblong, (1.5–)1.8–2.4 × (1–)1.2–1.4 mm, apex acute to ± acute, mucronate.

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

Flowers

stamens 2;

anthers (0.4–)0.5–0.8 mm, connective not prolonged;

styles 0.6–1(–1.4) mm;

stigmas (1–)1.4–2 mm.

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

Achenes

light to dark brown, oblong, (0.8–)0.9–1.1(–1.2) × 0.4–0.5(–0.6) mm, base cuneate to ± stipelike, apex truncate, apiculate, surfaces punctate to minutely punctate with raised cells.

biconvex, flattened, or trigonous.

Cyperus polystachyos

Cyperus

Phenology Fruiting summer–early fall.
Habitat Shores, ditches, swales between dunes
Elevation 0–200 m (0–700 ft)
Distribution
from FNA
AL; DC; DE; FL; GA; KY; LA; MA; MD; ME; MS; NC; NJ; NY; OK; PA; SC; TX; VA; WV; Mexico; Central America; South America; Asia; Africa
[WildflowerSearch map]
[BONAP county map]
from USDA
Pantemperate and tropical
[BONAP county map]
Discussion

Cyperus polystachyos is polymorphic. Adequate understanding of the infraspecific variations of C. polystachyos requires a worldwide examination, which is beyond the scope of this study. The segregates, C. fugax and C. filicinus, are recognized here; they have been usually accepted by recent workers (e.g., M. L. Fernald 1950; H. A. Gleason and A. Cronquist 1991).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 600 (96 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Source FNA vol. 23, p. 160. FNA vol. 23, p. 141. Authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
Parent taxa Cyperaceae > Cyperus > subg. Pycreus Cyperaceae
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Synonyms C. brizaeus, C. polystachyos var. leptostachyus, Pycreus polystachyos
Name authority Rottbøll: Descr. Pl. Rar., 21. (1772) Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754)
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