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many-spike flatsedge

false flatsedge, false nutsedge, straw color cyperus, straw-color flatsedge, straw-colour flatsedge

Habit Herbs, perennial (annual?), cespitose. Herbs, perennial (often flowering first year), rhizomatous.
Culms

trigonous or ± terete, (3–)20–45(–70) cm × (0.8–) 1.2–2 mm, glabrous.

basally swollen and cormlike, trigonous, (1–)20–40(–90) cm × 1–6 mm, glabrous or few prickles subapically.

Leaves

(2–) 6–10(–16), flat to V-shaped, (2–) 10–30(–60) cm × 1–4 mm.

flat, (10–)20–40 cm × 1–4(–8) mm, margins and midribs glabrous or scabridulous.

Inflorescences

spike 1, loosely ovoid to densely turbinate, 12–40 × 10–25 mm;

rachis 3–10 mm;

rays 1–8(–12), 0.5–3(–6) cm; 2d order rays absent;

if rays absent, inflorescence single dense cluster of spikelets, 12–40 mm diam.;

bracts (3–)4–6(–7), horizontal to ascending at 45°, flanged V-shaped, (2–)5–15(–23) cm × 1–4 mm.

spikes 1–4, loosely ovoid to oblong-cylindric; (6–)10–28(–50) × 10–40 mm;

rays sometimes absent, usually (1–)3–6(–8), 0.5–7(–25) cm;

if absent, inflorescence a congested head; 2d order rays sometimes present, 1–4(–9) cm;

rachis 8–18(–25) mm;

bracts (3–)5–7(–10), ascending at 30–45°, cross ribs sometimes conspicuous, (1–)10–30(–45) cm × (1–)2–4(–12) mm;

rachilla ± deciduous, wings 0.4–0.5 mm wide.

Spikelets

(5–)15–30(–40), spreading to appressed-ascending, linear-lanceoloid to linear, compressed, 8–18(–40) × (1–)1.3–1.6(–2) mm;

floral scales (8–)12–25(–66), laterally stramineous to light brown, medially green, laterally ribless, medially 3–5-ribbed, oblong, (1.5–)1.8–2.4 × (1–)1.2–1.4 mm, apex acute to ± acute, mucronate.

(5–)12–50, linear-lanceoloid, compressed, 5–30 × 0.6–0.9 mm;

floral scales ± deciduous, 3–11, appressed, medially green, laterally stramineous to pale brown, 3–4-ribbed laterally, (1–)3-ribbed medially, oblong-obovate, 3.2–4.5(–6) × 1–1.8 mm, apex ± acute, overlapping lower 1/4–1/2 of next scale.

Flowers

stamens 2;

anthers (0.4–)0.5–0.8 mm, connective not prolonged;

styles 0.6–1(–1.4) mm;

stigmas (1–)1.4–2 mm.

anthers 0.3–0.5 mm;

styles 1–2 mm;

stigmas 3–4 mm.

Achenes

light to dark brown, oblong, (0.8–)0.9–1.1(–1.2) × 0.4–0.5(–0.6) mm, base cuneate to ± stipelike, apex truncate, apiculate, surfaces punctate to minutely punctate with raised cells.

purplish brown, ± stipitate, narrowly oblong, (1.5–)1.8–2.4 × 0.5–0.6 mm, apex apiculate, acute, surfaces puncticulate.

Cyperus polystachyos

Cyperus strigosus

Phenology Fruiting summer–early fall. Fruiting summer.
Habitat Shores, ditches, swales between dunes Pond shores, ditches, damp, disturbed soils, sometimes weeds of gardens or croplands
Elevation 0–200 m (0–700 ft) 0–1500 m (0–4900 ft)
Distribution
from FNA
AL; DC; DE; FL; GA; KY; LA; MA; MD; ME; MS; NC; NJ; NY; OK; PA; SC; TX; VA; WV; Mexico; Central America; South America; Asia; Africa
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from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; MB; ON; QC; SK
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Discussion

Cyperus polystachyos is polymorphic. Adequate understanding of the infraspecific variations of C. polystachyos requires a worldwide examination, which is beyond the scope of this study. The segregates, C. fugax and C. filicinus, are recognized here; they have been usually accepted by recent workers (e.g., M. L. Fernald 1950; H. A. Gleason and A. Cronquist 1991).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus strigosus is usually among the more common Cyperus species throughout its range. Small individuals flowering the first year may be difficult to distinguish from C. polystachyos, C. odoratus, C. erythrorhizos, and C. esculentus, which may be sympatric. Cyperus strigosus has trigonous achenes and three stigmas, unlike C. polystachyos; C. strigosus has flattened spikelets, unlike the subcylindric ones of C. odoratus; C. strigosus has floral scales usually 3 mm or more, unlike the smaller (1.2–1.5 mm) ones of C. erythrorhizos; C. strigosus has deciduous floral scales and a cormlike stem base with stolons, unlike C. esculentus.

Cyperus strigosus appears to be closely related to the neotropical C. camphoratus Liebmann; both species have deciduous floral scales and deciduous spikelets (G. C. Tucker 1994). Plants segregated as C. stenolepis cannot be distinguished consistently from C. strigosus on any single charcter, rather they appear to be merely large individuals of C. strigosus with long floral scales and frequently septate inflorescence bracts (M. L. Horvat 1941).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 160. FNA vol. 23, p. 184.
Parent taxa Cyperaceae > Cyperus > subg. Pycreus Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. brizaeus, C. polystachyos var. leptostachyus, Pycreus polystachyos C. hansenii, C. stenolepis, C. strigosus var. hansenii, C. strigosus var. stenolepis, Mariscus stenolepis, Mariscus strigosus
Name authority Rottbøll: Descr. Pl. Rar., 21. (1772) Linnaeus: Sp. Pl. 1: 47. (1753)
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