Cyperus polystachyos |
Cyperus echinatus |
|
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many-spike flatsedge |
globe flatsedge, teasel sedge |
|
Habit | Herbs, perennial (annual?), cespitose. | Herbs, perennial, single-stemmed to loosely cespitose. |
Culms | trigonous or ± terete, (3–)20–45(–70) cm × (0.8–) 1.2–2 mm, glabrous. |
basally cormlike, trigonous, (15–)30–100 cm × 0.5–3.5 mm, glabrous. |
Leaves | (2–) 6–10(–16), flat to V-shaped, (2–) 10–30(–60) cm × 1–4 mm. |
flat to V-shaped, 10–65 cm × 3–9 mm, adaxial surface, margins minutely scabridulous. |
Inflorescences | spike 1, loosely ovoid to densely turbinate, 12–40 × 10–25 mm; rachis 3–10 mm; rays 1–8(–12), 0.5–3(–6) cm; 2d order rays absent; if rays absent, inflorescence single dense cluster of spikelets, 12–40 mm diam.; bracts (3–)4–6(–7), horizontal to ascending at 45°, flanged V-shaped, (2–)5–15(–23) cm × 1–4 mm. |
spikes densely globose to globose-ovoid, 8–17 mm wide; rays 3–12, 2–12 cm, scaberous adaxially especially distally; rachis 4–8 mm; bracts (3–)4–7, ascending at 30(–45)°, flat, 5–35 cm × 2–9 mm; rachilla persistent, wings 0.5–0.7 mm wide. |
Spikelets | (5–)15–30(–40), spreading to appressed-ascending, linear-lanceoloid to linear, compressed, 8–18(–40) × (1–)1.3–1.6(–2) mm; floral scales (8–)12–25(–66), laterally stramineous to light brown, medially green, laterally ribless, medially 3–5-ribbed, oblong, (1.5–)1.8–2.4 × (1–)1.2–1.4 mm, apex acute to ± acute, mucronate. |
50–100, oblong-lanceoloid, ± terete-quadrangular, (3.5–)4–7 × 1–1.4 mm; distal spikelet spreading or ascending; floral scales persistent, 3–5, appressed, stramineous to brownish, 4-ribbed laterally, oblong-elliptic, 3.5–4.5 × 1–1.8 mm, membranous, apex entire or emarginate with mucro to 0.3 mm. |
Flowers | stamens 2; anthers (0.4–)0.5–0.8 mm, connective not prolonged; styles 0.6–1(–1.4) mm; stigmas (1–)1.4–2 mm. |
anthers 0.4–0.8 mm; styles 0.5–0.6 mm; stigmas 1 mm. |
Achenes | light to dark brown, oblong, (0.8–)0.9–1.1(–1.2) × 0.4–0.5(–0.6) mm, base cuneate to ± stipelike, apex truncate, apiculate, surfaces punctate to minutely punctate with raised cells. |
brown, ± stipitate, oblong, (1.5–)1.8–2.3 × 0.5–0.6(–0.7) mm (1/2 length of floral scales), apex obtuse, surfaces puncticulate. |
Cyperus polystachyos |
Cyperus echinatus |
|
Phenology | Fruiting summer–early fall. | Fruiting summer–early fall. |
Habitat | Shores, ditches, swales between dunes | Disturbed, sunny sites, in mesic places, well-drained soils |
Elevation | 0–200 m (0–700 ft) | 0–500 m (0–1600 ft) |
Distribution |
AL; DC; DE; FL; GA; KY; LA; MA; MD; ME; MS; NC; NJ; NY; OK; PA; SC; TX; VA; WV; Mexico; Central America; South America; Asia; Africa
|
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; West Indies
|
Discussion | Cyperus polystachyos is polymorphic. Adequate understanding of the infraspecific variations of C. polystachyos requires a worldwide examination, which is beyond the scope of this study. The segregates, C. fugax and C. filicinus, are recognized here; they have been usually accepted by recent workers (e.g., M. L. Fernald 1950; H. A. Gleason and A. Cronquist 1991). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The records for Rhode Island and Wisconsin are according to M. L. Horvat (1941); we have not seen specimens from those states. Cyperus echinatus is usually recognized by its tight, nearly spheric spikes; it may occasionally be hard to distinguish from C. croceus and C. retrorsus. Compared to C. retrorsus, C. echinatus has larger spikelets and longer floral scales, anthers, and achenes. In contrast to C. echinatus, C. croceus has looser spikes, shorter, broader, greenish or yellowish floral scales, shorter, more ovoid achenes, and shorter anthers. Furthermore, C. echinatus is predominantly an inland species of roadsides, pastures, and other disturbed ground; C. retrorsus is primarily a coastal species and occurs in drier, sandier sites. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 160. | FNA vol. 23, p. 182. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. brizaeus, C. polystachyos var. leptostachyus, Pycreus polystachyos | Scirpus echinatus, C. ovularis, C. ovularis var. americanus, C. ovularis var. sphaericus, C. ovularis var. wolfii, C. wolfii, Kyllinga ovularis, Mariscus ovularis |
Name authority | Rottbøll: Descr. Pl. Rar., 21. (1772) | (Linnaeus) Alph. Wood: Class-book Bot. ed. s.n.(b), 734. (1861) |
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