Cyperus pilosus |
Cyperus entrerianus |
|
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fuzzy flatsedge |
woodrush flatsedge |
|
Habit | Herbs, annual or perennial, cespitose; stolons slender, to 5 cm × 1.5 mm. | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. |
Culms | trigonous, 25–90 cm × 2 mm, hispidulous or glabrous on angles distally. |
1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
Leaves | flat, 10–35 cm × 3–10 mm. |
(1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
Inflorescences | spikes 1–4(–6), loosely oblong-ovoid, 2–3 × 1–2 cm; rays 3–10, 1–16 cm; 2d order rays to 3 cm; rachis hispidulous, prickles slender, 0.1–0.2 mm; bracts 3–5, ± ascending, flat, 5–35 cm × 0.5–5(–7) mm; rachilla persistent, wingless. |
heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
Spikelets | 15–40, linear-lanceoloid, compressed, 6–12 × 1.5–2 mm; floral scales deciduous, 8–16, light brown with clear border, laterally 2–3-ribbed, ovate, 1.8–2 × 1.2–1.6 mm; apex entire, emarginate, or mucronulate, mucro at most 0.3 mm. |
(1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
Flowers | anthers 0.4 mm; styles 0.6 mm; stigmas 1 mm. |
stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
Achenes | brown, sessile, broadly ellipsoid, 1–1.2 × 0.5 mm, apex obtuse, apiculate, surfaces puncticulate. |
brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
Cyperus pilosus |
Cyperus entrerianus |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Rice fields, wetlands, emergent muddy shores | Roadside ditches, marshes |
Elevation | 0–50 m (0–200 ft) | 0–100 m (0–300 ft) |
Distribution |
FL; LA; MS; SC; Asia [Introduced in North America]
|
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
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Discussion | Cyperus pilosus was naturalized in the flora from eastern Asia, apparently early in the twentieth century. The combination of hispidulous rachis and loosely oblong-ovoid spikes is sufficient to distinguish Cyperus pilosus from any others with deciduous floral scales. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 166. | FNA vol. 23. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | |
Name authority | Vahl: Enum. Pl. 2: 354. (1805) | Boeckeler: Flora 61: 139. (1878) |
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