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Egyptian paper-reed, papyrus

tidalmarsh flatsedge

Habit Herbs, perennial, rhizomatous, stout. Herbs, perennial, stoloniferous; stolons terminated by tubers.
Culms

roundly trigonous, 300–500 cm × 15–45 mm, glabrous.

trigonous, 50–100 cm × 5–8 mm, glabrous.

Leaves

bladeless.

3–6, V-shaped, 20–50 cm × 5–8(–10) mm.

Inflorescences

spikes loosely cylindric, 10–20 × 6–10 mm;

rays 40–100, drooping or arching, slender, 10–30 cm; 2d order rays 8–20 cm;

bracts 4–10, ± erect, V-shaped, 3–8 cm × 4–15 mm; 2d order bracts 2–5, (1.5–)4–16 cm × 0.5–2 mm;

rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide.

spikes not appearing lateral, rays 5–8, 2–7(–12) cm;

bracts 3–4, ± horizontal to ascending at 45°, V-shaped, 5–25 cm × 1.5–8 mm;

spikes 1–4, ovoid, 2–4 × 2–3 cm;

rachis 1–2.7 cm, smooth or hispidulous;

rachilla persistent, wingless or with hyaline wings scarcely 0.1 mm wide.

Spikelets

6–30, slightly compressed, linear, ± quadrangular, 6–10 × 0.8–1 mm;

floral scales 6–16, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate-elliptic, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse.

7–14, flattened, lanceoloid, 10–20 × 2–2.5 mm;

floral scales 10–30, laterally dark brown to reddish brown, medially greenish, laterally 5–9-ribbed, ovate-deltate;

anthers 3, 0.5 mm;

styles 0.5 mm;

stigmas 1.2–1.6 mm.

Flowers

anthers 0.8–1 mm (connectives prolonged beyond anther as red subulate appendage 0.2–0.5 mm, apex usually setose);

styles 0.2–0.4 mm;

stigmas 0.8–1.2 mm.

Achenes

pale brown, sessile, oblong, 0.8–1 × 0.4 mm, apex scarcely apiculate, surfaces puncticulate.

brown, sessile, ellipsoid, 1.4–1.6 × 1.4–1.5 mm, apex obtuse, apiculate.

Cyperus papyrus

Cyperus serotinus

Phenology Fruiting summer. Fruiting summer.
Habitat Stream banks, marshes Tidal marshes
Elevation 0–30 m (0–100 ft) 0 m (0 ft)
Distribution
from FNA
FL; s Europe; sw Asia; Africa [Introduced in North America]
[BONAP county map]
from FNA
DE; NJ; PA; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Cyperus papyrus is conspicuous in the field by its great height, leafless culm, and open, hemispheric inflorescence with drooping rays.

Ancient Egyptians, Greeks, and Romans prepared paper from the pith of the culms of Cyperus papyrus, which is abundant along the Nile River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 173. FNA vol. 23, p. 164.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae > Cyperus > subg. Juncellus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms Chlorocyperus serotinus, Juncellus serotinus
Name authority Linnaeus: Sp. Pl. 1: 47. (1753) Rottbøll: Descr. Pl. Rar., 18. (1772)
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