The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Egyptian paper-reed, papyrus

Alabama swamp flatsedge

Habit Herbs, perennial, rhizomatous, stout. Herbs, perennial, solitary or cespitose, rhizomatous, coarse.
Culms

roundly trigonous, 300–500 cm × 15–45 mm, glabrous.

trigonous, (30–)60–130 cm × 1–4(–9) mm, densely papillose, otherwise glabrous.

Leaves

bladeless.

with cross ribs prominent, V-shaped, 30–100 cm × 5–13(–20) mm, margins, keel incisive with brownish prickles, papillose throughout, margins and midribs harshly scabrid.

Inflorescences

spikes loosely cylindric, 10–20 × 6–10 mm;

rays 40–100, drooping or arching, slender, 10–30 cm; 2d order rays 8–20 cm;

bracts 4–10, ± erect, V-shaped, 3–8 cm × 4–15 mm; 2d order bracts 2–5, (1.5–)4–16 cm × 0.5–2 mm;

rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide.

spikes 3–7, densely oblong-cylindric to ± globose, 10–28(–35) × 8–12(–15) mm;

rays 5–12, 1–16 cm, densely papillose;

bracts 5–12, ascending at 30–75°, flat to V-shaped, 4–50(–90) cm × 1–15 mm;

rachilla deciduous, wings persistent, (0.2–)0.3–0.5(–0.6) mm.

Spikelets

6–30, slightly compressed, linear, ± quadrangular, 6–10 × 0.8–1 mm;

floral scales 6–16, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate-elliptic, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse.

20–80, oblong-ellipsoid, ± terete, slightly compressed, 3–7 × 1–2(–2.7) mm;

distal spikelets spreading or ascending;

floral scales (2–)4–7, appressed, clear to brownish, reddish streaked, medially green, 9–11-ribbed, ovate, (2–)2.5–3.3 × 1.2–2.3 mm, apex acute, mucronate.

Flowers

anthers 0.8–1 mm (connectives prolonged beyond anther as red subulate appendage 0.2–0.5 mm, apex usually setose);

styles 0.2–0.4 mm;

stigmas 0.8–1.2 mm.

anthers 0.6–0.8 mm, styles 0.5–1 mm;

stigmas 1–2.3 mm.

Achenes

pale brown, sessile, oblong, 0.8–1 × 0.4 mm, apex scarcely apiculate, surfaces puncticulate.

brown, slightly stipitate, obovoid to broadly ellipsoid, (1.2–)1.5–1.6(–1.7) × 0.6–0.8 mm, apex apiculate, surfaces puncticulate.

Cyperus papyrus

Cyperus ligularis

Phenology Fruiting summer. Fruiting throughout the year.
Habitat Stream banks, marshes Beaches, brackish marshes, mangrove swamps, disturbed soils, ditches, riverbanks, coastal croplands
Elevation 0–30 m (0–100 ft) 0 m (0 ft)
Distribution
from FNA
FL; s Europe; sw Asia; Africa [Introduced in North America]
[BONAP county map]
from FNA
FL; Mexico; Central America; South America; w Africa
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus papyrus is conspicuous in the field by its great height, leafless culm, and open, hemispheric inflorescence with drooping rays.

Ancient Egyptians, Greeks, and Romans prepared paper from the pith of the culms of Cyperus papyrus, which is abundant along the Nile River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The papillose culms, rays, and leaves of Cyperus ligularis distinguish it from all other North American species of Cyperus.

Reports of Cyperus ligularis from California (G. C. Tucker 1993b) were based on specimens of C. owanii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 173. FNA vol. 23, p. 190.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms Mariscus rufus
Name authority Linnaeus: Sp. Pl. 1: 47. (1753) Linnaeus: Syst. Nat. ed. 10, 2: 867. (1759)
Web links