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Egyptian paper-reed, papyrus

Habit Herbs, perennial, rhizomatous, stout.
Culms

roundly trigonous, 300–500 cm × 15–45 mm, glabrous.

Leaves

bladeless.

Inflorescences

spikes loosely cylindric, 10–20 × 6–10 mm;

rays 40–100, drooping or arching, slender, 10–30 cm; 2d order rays 8–20 cm;

bracts 4–10, ± erect, V-shaped, 3–8 cm × 4–15 mm; 2d order bracts 2–5, (1.5–)4–16 cm × 0.5–2 mm;

rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide.

Spikelets

6–30, slightly compressed, linear, ± quadrangular, 6–10 × 0.8–1 mm;

floral scales 6–16, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate-elliptic, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse.

Flowers

anthers 0.8–1 mm (connectives prolonged beyond anther as red subulate appendage 0.2–0.5 mm, apex usually setose);

styles 0.2–0.4 mm;

stigmas 0.8–1.2 mm.

Achenes

pale brown, sessile, oblong, 0.8–1 × 0.4 mm, apex scarcely apiculate, surfaces puncticulate.

Cyperus papyrus

Phenology Fruiting summer.
Habitat Stream banks, marshes
Elevation 0–30 m (0–100 ft)
Distribution
from FNA
FL; s Europe; sw Asia; Africa [Introduced in North America]
[BONAP county map]
Discussion

Cyperus papyrus is conspicuous in the field by its great height, leafless culm, and open, hemispheric inflorescence with drooping rays.

Ancient Egyptians, Greeks, and Romans prepared paper from the pith of the culms of Cyperus papyrus, which is abundant along the Nile River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 173.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Name authority Linnaeus: Sp. Pl. 1: 47. (1753)
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