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Owan's flatsedge

black flatsedge, brown cyperus

Habit Herbs, perennial, cespitose, rhizomatous. Herbs, perennial, rhizomatous.
Culms

roundly trigonous, 80–120 cm × 3–6 mm, glabrous.

solitary or close together, trigonous, 5–40(–95) cm, glabrous.

Leaves

5–12, V-shaped, 30–75 cm × 8–14 mm, septate, leathery, margins harshly scabrid.

(2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm.

Inflorescences

spikes 1–4 per ray, loosely ovoid to broadly ellipsoid, 12–20 × 8–12 mm;

rays 6–9, 3–10(–12) cm; 2d order rays 1–4, 1–2 cm;

rachis 4–12 mm;

bracts 5–10, horizontal to ascending at 30°, 15–40 cm × 4–8(–11) mm;

rachilla deciduous, wings persistent, hyaline, fragile, 0.3 mm wide, covering less than 1/2 of achene.

spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.;

rachis 1–2 mm, or absent;

rays 1–2, 3–45 mm;

if rays absent, inflorescence sessile, dense, 6–20 mm diam;

bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm.

Spikelets

40–80, narrowly ellipsoid to ellipsoid, roughly quadrangular to weakly flattened, 5–8 × 1.2–1.5 mm;

distal spikelet spreading or ascending;

floral scales (1–)2(–3), ascending, brick red, medially greenish, laterally 3–4-ribbed, medially 1–3-ribbed, green, ovate, 2.2–3.5 × 1.5–1.8 mm, apex subacute.

(3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm;

floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse.

Flowers

anthers (0.7–)1–1.5 mm, filaments 4–4.7 mm;

styles 0.5–0.9 mm;

stigmas 2–3 mm.

stamens 2;

anthers 0.6–0.8 mm, connectives not prolonged;

styles 0.4–0.6 mm;

stigmas 0.7–2.1 mm.

Achenes

brown, ellipsoid to oblong-fusiform, 1.7–1.8 × 0.7 mm, base cuneate, apex subacute and apiculate, surfaces finely puncticulate to glabrous.

brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate.

Cyperus owanii

Cyperus niger

Phenology Fruiting spring–fall. Fruiting summer.
Habitat Disturbed, damp thickets, open woods Swamps, ditches, wet pastures, seeps in montane forests
Elevation 0–200 m (0–700 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
CA; s Africa [Introduced in North America]
[BONAP county map]
from FNA
AZ; CA; NM; OK; TX; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus owanii is easily distinguished from other sedges in California by the large size, reddish spikelets, and secondary rays. It was first collected in California in 1938.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 184. FNA vol. 23.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae > Cyperus > subg. Pycreus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms Mariscus umbilensis C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus
Name authority Boeckeler: Flora 61: 29. (1878) Ruiz & Pavón: Fl. Peruv. 1: 47. (1798)
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