Cyperus niger |
Cyperus tetragonus |
|
---|---|---|
black flatsedge, brown cyperus |
fourangle flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, cespitose, rhizomatous. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
trigonous, (20–) 30–80 cm × 1–2.5 mm, glabrous or occasionally with scattered prickles on distal angles. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
9–11, flat to slightly V-shaped, 25–80 cm × 3.5–10 mm. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
spikes (1–)3–6, loosely cylindric or oblong with parallel sides, 1.5–4 cm × 10–17 mm; rays 6–12, 2–25 cm, glabrous or rarely scabridulous proximal to spikes; bracts 6–11, ascending at 45°, flat, (1–)10–20(–26) cm × 1–10 mm; rachilla deciduous, wings persistent, 0.4–0.5 mm wide. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
10–60, narrowly ellipsoid, quadrangular, 5–8 × 1.5–2 mm; floral scales persistent, (2–)3–7, appressed, clear at maturity, laterally reddish to purplish brown, medially greenish, often erose at maturity, laterally 3–5-ribbed, medially 3-ribbed, ovate-elliptic, 2.5–3.2 × 1.8–2.1 mm, apex obtuse. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
anthers 0.8–1 mm; styles 0.3–0.5 mm; stigmas 2–2.5 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
brown to reddish brown, sessile, oblong-ovoid to broadly ellipsoid, slightly dorsiventrally flattened, 1.8–2 × 0.7–0.8 mm, apex obtuse, apiculate, surfaces papillose to puncticulate. |
Cyperus niger |
Cyperus tetragonus |
|
Phenology | Fruiting summer. | Fruiting summer–fall (Jun–Nov). |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Open woods, thickets, barrier islands |
Elevation | 0–1200 m (0–3900 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
AL; AZ; FL; GA; MS; NC; NM
|
Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The status of Cyperus pringlei from the southwestern United States requires further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 189. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | C. pringlei, C. tetragonus var. pringlei, Mariscus tetragonus |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | Elliott: Sketch Bot. S. Carolina 1: 71. (1821) |
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