Cyperus niger |
Cyperus ligularis |
|
---|---|---|
black flatsedge, brown cyperus |
Alabama swamp flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, solitary or cespitose, rhizomatous, coarse. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
trigonous, (30–)60–130 cm × 1–4(–9) mm, densely papillose, otherwise glabrous. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
with cross ribs prominent, V-shaped, 30–100 cm × 5–13(–20) mm, margins, keel incisive with brownish prickles, papillose throughout, margins and midribs harshly scabrid. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
spikes 3–7, densely oblong-cylindric to ± globose, 10–28(–35) × 8–12(–15) mm; rays 5–12, 1–16 cm, densely papillose; bracts 5–12, ascending at 30–75°, flat to V-shaped, 4–50(–90) cm × 1–15 mm; rachilla deciduous, wings persistent, (0.2–)0.3–0.5(–0.6) mm. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
20–80, oblong-ellipsoid, ± terete, slightly compressed, 3–7 × 1–2(–2.7) mm; distal spikelets spreading or ascending; floral scales (2–)4–7, appressed, clear to brownish, reddish streaked, medially green, 9–11-ribbed, ovate, (2–)2.5–3.3 × 1.2–2.3 mm, apex acute, mucronate. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
anthers 0.6–0.8 mm, styles 0.5–1 mm; stigmas 1–2.3 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
brown, slightly stipitate, obovoid to broadly ellipsoid, (1.2–)1.5–1.6(–1.7) × 0.6–0.8 mm, apex apiculate, surfaces puncticulate. |
Cyperus niger |
Cyperus ligularis |
|
Phenology | Fruiting summer. | Fruiting throughout the year. |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Beaches, brackish marshes, mangrove swamps, disturbed soils, ditches, riverbanks, coastal croplands |
Elevation | 0–1200 m (0–3900 ft) | 0 m (0 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
FL; Mexico; Central America; South America; w Africa
|
Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The papillose culms, rays, and leaves of Cyperus ligularis distinguish it from all other North American species of Cyperus. Reports of Cyperus ligularis from California (G. C. Tucker 1993b) were based on specimens of C. owanii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 190. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | Mariscus rufus |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | Linnaeus: Syst. Nat. ed. 10, 2: 867. (1759) |
Web links |